The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
In the Theraphosae the coxa resembles that of the ambulatory leg, but in
other spiders it is furnished, on the inner side, with a blade-like
projection, the “maxilla” (Fig. 178). The shape of the maxillae and the
degree of their inclination towards the labium are of considerable
taxonomic importance. The inner border of the maxilla is furnished with
a tuft of hairs, which assist in retaining the juices expressed by the
chelicerae, and its anterior border presents a cutting edge with a
finely dentated ridge called the “serrula.”
In the female, and in the immature male, the remaining joints differ
little from those of the legs, except that the tarsal joint is either
clawless or has a single claw, which is generally smooth, and is never
much dentated.
At the last moult but one the male pedipalp appears tumid at the end,
and after the last moult the tarsus is seen to have developed a
remarkable copulatory apparatus, the “palpal organ,” comparatively
simple in some families, but in others presenting an extraordinary
complexity of structure.
[Illustration:
FIG. 179.—Diagram of palpal organ. 1, Tarsus; 2, bulb; 3, receptaculum
seminis; 4, its aperture; 5, style; 6, haematodocha; 7, alveolus; 8,
tibia.
]
=Palpal Organs.=—Externally the essential parts of the palpal organ are
three, the “haematodocha,” the “bulb,” and the “style.” The spines and
projections, or “apophyses,” which often accompany the palpal organ
proper, are of secondary importance, and in many spiders are entirely
absent; nor is their function when present at all clear; but the
infinite variety of design which they exhibit, and their singular
uniformity in all the males of a species, render them of the utmost
value as specific characteristics.
The “haematodocha” is the portion of the palpal organ attached to the
tarsus, and often received into an excavation, the “alveolus,” on its
under surface. It is a fibro-elastic bag, in its normal collapsed state
usually somewhat spirally disposed round the base of the following
portion, the “bulb.”
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