The Cambridge natural history, Vol. 04 (of 10)Warburton, Cecil
Science
The Cambridge natural history, Vol. 04 (of 10)
Warburton, Cecil
Animals
=The Stomodaeum= consists of the pharynx, the oesophagus, and the
sucking stomach. As we have said, the mouth is to be found between the
rostrum and the labium. It opens into the pharynx, the anterior wall of
which is formed by a chitinous plate on the inner surface of the
rostrum, sometimes called the palate. As the inner surfaces of the
rostrum and labium are practically flat, the cavity of the pharynx would
be obliterated when they are pressed together, were it not for a groove
running down the centre of the palate, which the apposed labium converts
into a tube, up which the fluids of the prey are sucked. In the
Theraphosidae there is a corresponding groove on the inner surface of
the labium.
At the top of the pharynx, which is nearly perpendicular, the canal
continues backwards and upwards as a narrow tube, the oesophagus,
passing right through the nerve-mass, which embraces it closely on all
sides, to the sucking stomach. At the commencement of the oesophagus is
the opening of a gland, probably salivary, which is situated in the
rostrum.
[Illustration:
FIG. 185.—Diagram showing the anatomy of the cephalothorax of a
Spider. The right alimentary diverticulum has been removed. _a_,
Aorta; _c_, left diverticulum with secondary caeca; _e_,
endosternite; _oes_, oesophagus, descending to the mouth; _s_,
sucking stomach; _sh_, dorsal shield of sucking stomach.
]
We now reach the sucking stomach, which occupies the centre of the
cephalothorax. It is placed directly over a skeletal plate, the
“endosternite” (Fig. 185, _e_), to which its lower surface is connected
by powerful muscles, while its upper wall is protected by a hard plate
or “buckler,” which is similarly attached to the roof of the
cephalothorax in the region of the “fovea media.” The walls of the
stomach are not themselves muscular, but by the contraction of the
muscles above mentioned its cavity is enlarged, and fluids from the
pharynx are pumped up into it.
The canal thus far is lined by chitin, like the exterior of the body,
and forms a sort of complicated mouth-apparatus.
The =Mesenteron= lies partly in the cephalothorax and partly in the
abdomen. The thoracic portion, shortly behind the sucking stomach, sends
forward on either side a large branch or “diverticulum,” from each of
which five secondary branches or “caeca” are given off (Fig. 185). Of
these the anterior pair sometimes join, thus forming a complete ring;
but usually, though adjacent, they remain distinct. The other four pairs
of caeca curve downwards, protruding into the coxae of the legs, where
they often terminate, but sometimes (_Epeira_) they continue their curve
until they meet, though they never fuse, under the nerve-mass. Behind
the origin of the diverticula the mesenteron continues as a widish tube,
and shortly passes through the pedicle and enters the abdomen, where,
curving slightly upwards, it proceeds along the middle line till it ends
in the proctodaeum.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account