The Cambridge natural history, Vol. 05 (of 10)Sedgwick, Adam
Science
The Cambridge natural history, Vol. 05 (of 10)
Sedgwick, Adam
Animals
The chief characters on which Brauer bases his system are: (1) The
existence or absence of wings. (2) The condition of the mouth, and whether
it undergoes radical changes in the ontogeny, arriving thus at the
categories Menognatha, Metagnatha, and Menorhyncha, as we have mentioned on
p. 161. (3) The metamorphosis; the grouping adopted being Ametabola,
Hemimetabola, Metabola. (4) The number of the Malpighian tubules;
Oligonephria, Polynephria. (5) The nature of the wings, the relative
proportions of the thoracic segments, and some other characters.
Brauer's treatise is accompanied by a valuable and in many respects very
sagacious consideration of the generalised characters of the Insecta; as a
classification based partly on generalisations and partly on structures, it
is, so far as the present {176}condition of our knowledge goes, a good one.
But it is of little value as a practical guide, and as a basis for
theoretical speculation it cannot be treated as of importance, because the
generalisations it makes use of are premature, owing to the small
proportion of the forms that have been examined. And even now the groups
adopted are known to be subject to many exceptions.
Thus it begins by a division of Insecta into winged and wingless; but the
winged division is made to comprehend an enormous number of wingless
Insects, whole subdivisions of Orders such as the Mallophaga being placed
in the winged series, although all are without wings. This first division
is indeed entirely theoretical; and if a classification on generalisations
were adopted, it would be more natural to begin with the old division into
Homomorpha and Heteromorpha, and treat the Order Aptera as the first
division of the Homomorpha, while the Heteromorpha would commence with the
Ephemeridae and Odonata, in which, though the individual in the early part
of the ontogeny is very different from the perfect Insect, there is no
marked division of the later larval and the pupal stages. Brauer's system
is also defective inasmuch as it takes no account of the embryological or
oogenetic processes, though these are of equal importance with the later
phases of the Ontogeny. Even as regards the division into Orders, it is far
from being free from reproach; for instance, the separation of the
Dermaptera from the Orthoptera, while Rhynchota remains intact, although
including a more extensive series of heterogeneous forms; the division of
the Neuroptera into widely separated groups, each of which is treated as
equivalent to the great Orders, such as Coleoptera (in which Strepsiptera
are included), Hymenoptera, and Diptera, is not reasonable. The association
of Mallophaga and Termitidae, while Dermaptera are separated from
Orthoptera, is also undeniably arbitrary, and other similar disparities are
to be seen on scrutinising the details of the system.
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