The Cambridge natural history, Vol. 05 (of 10)Sedgwick, Adam
Science
The Cambridge natural history, Vol. 05 (of 10)
Sedgwick, Adam
Animals
however, the forceps, and this is the most marked collective character of
the group. These curious organs exhibit a very great variety; in some cases
they are as long as the whole of the rest of the body, in others they are
provided with tynes; sometimes they are quite asymmetrical, as in
_Anisolabis tasmanica_ (Fig. 113); in _Opisthocosmia cervipyga_, and many
others they are curiously distorted in a variety of ways. The
classification of the earwigs is still in a rudimentary state; the number
of joints in the antennae, the form of the feet, and (in the terrestrial
forms) the shape of the rudimentary wing-cases and wings being the
characters that have been made most use of by {216}systematists; no
arrangement into sub-families or groups of greater importance than genera
is adopted.
The only particulars we have as to the embryological development of the
earwig are due to Heymons.[144] The forceps spring from the eleventh
abdominal segment, and represent the cerci of other Orthoptera. An
egg-tooth is found to be present on the head for piercing the egg-shell.
The embryo reverses its curved position during the development, as other
Orthoptera have been observed to do, but in a somewhat different manner,
analogous to that of the Myriapods.
Several fossil Forficulidae are known; specimens belonging to a peculiar
genus have been described from the Lower Lias of Aargau and from the
Jurassic strata in Eastern Siberia, but the examples apparently are not in
a very satisfactory state of preservation. In the Tertiary formations
earwigs have been found more frequently. Scudder has described eleven
species of one peculiar genus from the Lower Miocene beds at Florissant in
Colorado; some of these specimens have been found with the wings expanded,
and no doubt that they were fully developed Forficulidae can exist. The
fossil species of earwigs as yet known do not display so remarkable a
development of the forceps as existing forms do.
[Illustration: FIG. 113.—_Anisolabis tasmanica_ ♂.]
Brauer and others treat the Forficulidae as a separate Order of
Insects—Dermaptera—but the only structural characters that can be pointed
out as special to the group are the peculiar form of the tegmina and hind
wings—which latter, as we have said on p. 206, are considered by some to be
formed on essentially the same plan as those of other Orthoptera—the
imbrication of the segments, and the forceps terminating the body. The
development, so far as it is known, is that of the normal Orthoptera. Thus
the Forficulidae are a very distinct division of Orthoptera, the characters
that separate them being comparatively slight, though there are no
intermediate forms. Some of those who treat the Dermaptera as a sub-Order
equivalent to the rest of the divisions of the Order, call the latter
combination Euorthoptera.
{217}FAM. II. HEMIMERIDAE.
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