The Cambridge natural history, Vol. 05 (of 10)Sedgwick, Adam
Science
The Cambridge natural history, Vol. 05 (of 10)
Sedgwick, Adam
Animals
The head is most variable in size and form; as a part of its surface is
occupied by the eyes and as these organs differ in shape, extent, and
position to a surprising degree, it is not a matter for astonishment that
it is almost impossible to agree as to terms for the areas of the head. Of
the sclerites of the head itself there are only three that are sufficiently
constant and definite to be worthy of description here. These are the
clypeus, the epicranium, and the gula. The clypeus is situate on the upper
surface of the head-capsule, in front; it bears the labrum which may be
briefly described as a sort of flap forming an upper lip. The labrum is
usually possessed of some amount of mobility. The clypeus itself is
excessively variable in size and form, and sometimes cannot be delimited
owing to the obliteration of the suture of connexion with the more
posterior part of the head; it is rarely or never a paired piece.
Occasionally there is a more or less distinct piece interposed between the
clypeus and the labrum, and which is the source of considerable difficulty,
as it may be taken for the clypeus. Some authors call the clypeus the
epistome, but it is better to use this latter term for the purpose of
indicating the part that is immediately behind the labrum, whether that
part be the clypeus, or some other sclerite; the {93}term is very
convenient in those cases where the structure cannot be, or has not been,
satisfactorily determined morphologically.
In Figure 50 the parts usually visible on the anterior aspect of the head
and its appendages are shown so far as these latter can be seen when the
mouth is closed; in the case of the Insect here represented the bases of
the mandibles are clearly seen (_g_), while their apical portions are
entirely covered by the labrum, just below the lower margin of which the
tips of the maxillae are seen, looking as if they were the continuations of
the mandibles.
The labrum is a somewhat perplexing piece, morphologists being not yet
agreed as to its nature; it is usually placed quite on the front of the
head, and varies extremely in form; it is nearly always a single or
unpaired piece; the French morphologist Chatin considers that it is really
a paired structure.
[Illustration: FIG. 50.—Front view of head of field-cricket (_Gryllus_):
_a_, epicranium; _b_, compound eye; _c_, antenna; _d_, post-: _e_,
ante-clypeus; _f_, labrum; _g_, base of mandible; _h_, maxillary palpus;
_i_, labial palpus; _k_, apex of maxilla.]
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