The Cambridge natural history, Vol. 05 (of 10)Sedgwick, Adam
Science
The Cambridge natural history, Vol. 05 (of 10)
Sedgwick, Adam
Animals
There is another part of the mouth to which we may call special attention,
as it has recently attracted more attention than it formerly did; it is a
membranous lobe in the interior of the mouth, very conspicuous in
Orthoptera, and called the tongue, lingua, or hypopharynx; it reposes, in
the interior of the mouth (Fig. 51, _o_), on the middle parts of the front
of the labium; it is probably not entirely lost in Coleoptera, but enters
into the composition of the {97}complex middle part of the lip by
amalgamation with the paraglossae. It has recently been proposed to treat
this lingua as the morphological equivalent of the labium or of the
maxillae, giving it the name of the endolabium, but the propriety of this
course remains to be proved;[20] the view is apparently suggested chiefly
by the structure of the mouth of _Hemimerus_, a very rare and most peculiar
Insect that has not as yet been sufficiently studied.
As the maxillae and labium are largely used by taxonomists in the
systematic arrangement of the mandibulate Insects, we give a figure of them
as seen in Coleoptera, where the parts, though closely amalgamated, can
nevertheless be distinguished. This Fig. 52 should be compared with Fig.
51.
In speaking of the segments of the body we pointed out that they were not
separate parts but constituted an uninterrupted whole, and it is well to
remark here that this is also true of the gnathites. Although the mouth
parts are spoken of as separate pieces, they really form only projections
from the great body wall. Fig. 51, B, shows the intimate connexion that
exists between the maxillae and labium; the continuity of the mandibles
with the membrane of the buccal cavity is capable of very easy
demonstration.
The head bears, besides the pieces we have considered, a pair of antennae.
These organs, though varying excessively in form, are always present in the
adult Insect, and exist even in the majority of young Insects. They are
very mobile, highly sensitive organs, situate on or near the front part of
the head. The antennae arise in the embryo from the procephalic lobes, the
morphological import of which parts is one of the most difficult points
connected with Insect embryology.
The eyes of Insects are of two sorts, simple and compound. The simple eyes,
or ocelli, vary in number from one to as many as eighteen or twenty; when
thus numerous they are situated in groups on each side of the head. In
their most perfect form, as found in adult aculeate Hymenoptera, in
Orthoptera and Diptera, ocelli are usually two or three in number, and
present the appearance of small, perfectly transparent lenses inserted in
the integument. In their simplest form they are said to consist of some
masses of pigment in connexion with a nerve.
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