The Cambridge natural history, Vol. 05 (of 10)Sedgwick, Adam
Science
The Cambridge natural history, Vol. 05 (of 10)
Sedgwick, Adam
Animals
is conspicuously the more anterior, while in others the epimeron is placed
much above it, and may extend nearly as far forwards as it. It may be said,
as a rule, that when the sternum extends farther backwards than the notum,
the epimeron is above the episternum, as in many Coleoptera; but if the
sternum be anterior to the notum, then the episternum is superior to the
epimeron, as in dragon-flies. We would here again reiterate the fact that
these "pieces" are really not separate parts, but are more or less
indurated portions of a continuous integument, which is frequently entirely
occupied by them; hence a portion of a sclerite that in one species is
hard, may in an allied form be wholly or partly membranous, and in such
case its delimitation may be very evident on some of its sides, and quite
obscure on another.
[Illustration: FIG. 55.—Mesothorax of _Dytiscus_, after Audouin. A, notum;
A', pieces of the notum separated: _a_, praescutum; _b_, scutum; _c_,
scutellum; _d_, post-scutellum: B, the sternum and pleura united; B', their
parts separated: _a_, sternum; _c_, episternum; _d_, parapteron; _e_,
epimeron.]
{102}The parapteron of Audouin does not appear to be really a distinct
portion of the pleuron; in the case of _Dytiscus_ it is apparently merely a
thickening of an edge. Audouin supposed this part to be specially connected
with the wing-articulation, and the term has been subsequently used by
other writers in connexion with several little pieces that exist in the
pleural region of winged Insects.
The prothorax is even more subject to variation in its development than the
other divisions of the thorax are. In the Hymenoptera the prosternum is
disconnected from the pronotum and is capable, together with the first pair
of legs, of movement independent of its corresponding dorsal part, the
pronotum, which in this Order is always more or less completely united with
the meso-thorax; in the Diptera the rule is that the three thoracic
segments are closely consolidated into one mass. In the majority of Insects
the prothorax is comparatively free, that is to say, it is not so closely
united with the other two thoracic segments as they are with one another.
The three thoracic rings are seen in a comparatively uniform state of
development in a great number of larvae; also in the adult stages of some
Aptera, and among winged insects in some Neuroptera such as the Embiidae,
Termitidae, and Perlidae. In Lepidoptera the pronotum bears a pair of
erectile processes called patagia; though frequently of moderately large
size, they escape observation, being covered with scales and usually
closely adpressed to the sides of the pronotum.
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