The Cambridge natural history, Vol. 06 (of 10)Sharp, David
Science
The Cambridge natural history, Vol. 06 (of 10)
Sharp, David
Animals
In Britain we have only two representatives of the sub-family, viz. _Ponera
contracta_, a small ant of dirty-yellow colour, found rarely in the
Southern counties, living in moss or under stones. Its colonies consist of
only a few individuals; Forel giving fifty as the highest number he has
observed. The second species, _P. punctatissima_, presents the almost
unique peculiarity of possessing two forms of the male sex, one of them
resembling the worker in most of its peculiarities, and in being destitute
of wings, while the other is winged, as is usual in male ants. In the
island of St. Vincent another species of _Ponera_ has been discovered
having an apterous and worker-like male, and was named by Forel _P.
ergatandria_.[71] The discovery of this form has led him to express some
doubt as to whether _Ponera punctatissima_ has two forms of males; but it
seems probable that it really is so, the ergatoid males being produced
under somewhat different circumstances from the normal males. We shall
subsequently see that _Cardiocondyla_ and a few other Myrmicides exhibit an
analogous peculiarity.
The genus _Myrmecia_ is confined to the Australian continent and Tasmania,
and includes a considerable number of species of large and moderate-sized
ants, the classification of which has been a subject of difference of
opinion. This has arisen from the fact that the nodes of the abdominal
pedicel are more similar to those existing in the Myrmicides than to those
of the typical Ponerides. There are, however, some American members of the
latter sub-family (_Paraponera clavata_, _e.g._) that differ but little in
this point from _Myrmecia_, and, moreover, the pupae of _Myrmecia_ are
enclosed in a cocoon, while in the Myrmicides they are usually naked. On
the other hand the nests are, it appears, very large and populous, more
like what exists in the Myrmicides; there is no true stridulating organ on
the first abdominal segment. The genus is therefore one of those
interesting anomalies that form so large a proportion of the Australian
{173}fauna, and will probably be ultimately treated as a distinct
sub-family. There are about thirty species.
The ants of this genus are well known to the residents in Australia, where
they are called "bull-dog ants." They form large mounds of earth for their
nests. The workers, and females (Fig. 76) are much alike except during the
period when the latter are still carrying their wings. The males, however,
differ considerably, being of more slender form, and possessing only
insignificant mandibles, and straight antennae with a quite short basal
joint.
[Illustration: Fig. 76—_Myrmecia pyriformis._ Australia. Female after
casting wings.]
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