The Cambridge natural history, Vol. 06 (of 10)Sharp, David
Science
The Cambridge natural history, Vol. 06 (of 10)
Sharp, David
Animals
CLASSIFICATION.—Owing to the hardness of the skeleton, beetles shrivel but
little after death, so that the form and relations of the various sclerites
can usually be detected with ease. These sclerites seem to be remarkably
constant (except in the case of sexual distinctions) within the limits of
each species, and are very useful for the formation of genera and groups of
genera; but they vary so much outside the limits mentioned that it is very
difficult to make use of them for defining the larger groups. Hence it is
not easy to frame accurate definitions of the families, and still less so
to arrange these families in more comprehensive series. The natural
difficulty has been much increased by the habit coleopterists have of
framing their definitions on what is visible without the aid of dissection.
Nevertheless considerable progress has been made. We are obliged at present
to adopt upwards of eighty families; and we are able to distinguish on
positive characters five series; this leaves a large number of forms still
unclassified, and these we have here associated as a sixth series, which we
have called Coleoptera Polymorpha. This series corresponds with the two
series called in books Clavicornia and Serricornia. As it is admitted to be
impossible to define these two series, we think it much better to act
accordingly, and to establish for the present a great group that can only
be characterised by the fact that its members do not belong to any of the
other five series. No doubt a larger knowledge of development, coupled with
the advance of comparative anatomy, will ultimately bring about a better
state of affairs. The Strepsiptera, with one family Stylopidae, are only
provisionally included among the Coleoptera. These six series are fairly
equal as regards extent. Though the Polymorpha includes the larger number
of forms, yet a large part of them belong to four great families
(Staphylinidae, Buprestidae, Elateridae, Malacodermidae), which are easily
recognisable, so that the number of unmanageable forms is not really great.
Indeed, an acquaintance with the external anatomy of two or three dozen
species, selected as typical, would enable a student to classify {190}with
tolerable certainty the vast majority of species that he would subsequently
meet with.
Series 1. _Lamellicornia._—Antennae with the terminal joints leaf-like
(or broader than the others, if not actually leaf-like), and capable of
separation and of accurate apposition. Tarsi five-jointed.
Series 2. _Adephaga_—(_Caraboidea_ of some authors).—Antennae never
lamelliform, thin at the end; all the tarsi five-jointed, with the fourth
joint quite distinct. Maxillae highly developed, with the outer lobe
slender and divided into two segments so as to be palpiform. Abdomen with
six (or more) ventral segments visible.
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