The Cambridge natural history, Vol. 06 (of 10)Sharp, David
Science
The Cambridge natural history, Vol. 06 (of 10)
Sharp, David
Animals
There are a considerable number of blind members of this family: some of
them live in caverns; these belong chiefly to the genus _Anophthalmus_,
species of which have been detected in the caves of the Pyrenees, of
Austria, and of North America. It has been shown that the optic nerves and
lobes, as well as the external organs of vision, are entirely wanting in
some of these cave Carabidae; the tactile setae have, however, a larger
development than usual, and the Insects are as skilful in running as if
they possessed eyes. _Anophthalmus_ is closely related to our British genus
_Trechus_, the species of which are very much given to living in deep
crevices in the earth, or under large stones, and have some of them very
small eyes. In addition to these cavernicolous _Anophthalmus_, other blind
Carabidae have been discovered during recent years in various parts of the
world, where they live under great stones deeply embedded in the earth;
these blind lapidicolous Carabidae are of extremely minute size and of most
sluggish habits; the situations in which they are found suggest that many
successive generations are probably passed under the same stone. Not a
single specimen has ever been found above ground. The minute {206}Carabids
of the genus _Aëpus_, that pass a large part of their lives under stones
below high-water mark (emerging only when the tide uncovers them), on the
borders of the English Channel and elsewhere, are very closely allied to
these blind Insects, and have themselves only very small eyes, which,
moreover, according to Hammond and Miall, are covered in larger part by a
peculiar shield.[87] A few Carabidae, of the genera _Glyptus_ and
_Orthogonius_, are believed to live in the nests of Termites. Savage found
the larva of _G. sculptilis_ in the nests of _Termes bellicosus_; it has
been described by Horn, and is said to bear so great a resemblance to young
queens of the Termites as to have been mistaken for them.[88] Mr. Haviland
found _Rhopalomelus angusticollis_ in Termites' nests in South Africa.
Péringuey states that it emits a very strong and disagreeable odour. It is
probable that it preys on the Termites, and this also is believed to be the
habit of the Ceylonese _Helluodes taprobanae_. Some species of the
Mediterranean genus _Siagona_ stridulate by means of a file on the under
surface of the prothorax, rubbed by a striate area, adapted in form, on the
anterior femora.
A valuable memoir on the classification of this important family is due to
the late Dr. G. H. Horn;[89] he arranges Carabidae in three sub-families;
we think it necessary to add a fourth for _Mormolyce_:
1. Middle coxal cavities enclosed externally by the junction of the meso-
and meta-sternum; neither epimeron nor episternum attaining the
cavity.
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