The Cambridge natural history, Vol. 06 (of 10)Sharp, David
Science
The Cambridge natural history, Vol. 06 (of 10)
Sharp, David
Animals
FAM. 60. ELATERIDAE (_Click-beetles_).—_Antennae more or less serrate along
the inner margin, frequently pectinate, rarely filiform. Front coxae small,
spherical. Thorax usually with hind angles more or less prolonged
backwards; with a prosternal process that can be received in, and usually
can move in, a mesosternal cavity. Hind coxa with a plate, above which the
femur can be received. Visible ventral segments usually five, only the
terminal one being mobile. Tarsi five-jointed._ This large family of
Coleoptera comprises about 7000 species. Most of them are readily known by
their peculiar shape, and by their faculty of resting on the back,
stretching themselves out flat, and then suddenly going off with a click,
and thus jerking themselves into the air. Some, however, do not possess
this faculty, and certain of these are extremely difficult to recognise
from a definition of the family. According to Bertkau[133] our British
_Lacon murinus_ is provided near the tip of the upper side of the abdomen
with a pair of eversible glands, comparable with those that are better
known in Lepidopterous larvae. He states that this Insect does not try to
escape by leaping, but shams death and "stinks away" its enemy. The glands,
it would appear, become exhausted after the operation has been repeated
many times. The extent of the leap executed by click-beetles differs
greatly; in some species it is very slight, and only just sufficient to
turn the Insect right side up when it has been placed on its back. In some
cases the Insects go through the clicking movements with little or no
appreciable result in the way of consequent propulsion. Although it is
difficult to look on this clicking power as of very great value to the
Elateridae, yet their organisation is profoundly modified so as to permit
its accomplishment. The junction of the prothorax with the after-body
involves a large number of pieces which are all more or less changed, so
that the joint is endowed with greater mobility than usual; while in the
position of repose, on the other hand, the two parts are firmly locked
together. The thoracic stigma is of a highly remarkable nature, and the
extensive {258}membrane in which it is placed appears to be elastic.
Although the mechanics of the act of leaping are still obscure, yet certain
points are clear; the prosternal process possesses a projection, or notch,
on its upper surface near the tip; as a preliminary to leaping, this
projection catches against the edge of the mesosternal cavity, and as long
as this position is maintained the Insect is quiescent; suddenly, however,
the projection slips over the catch, and the prosternal process is driven
with force and rapidity into the mesosternal cavity pressing against the
front wall thereof, and so giving rise to the leap.
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