The Cambridge natural history, Vol. 07 (of 10)Boulenger, George Albert
Science
The Cambridge natural history, Vol. 07 (of 10)
Boulenger, George Albert
Animals
As regards their fixation, the more primitive forms of teeth, such as those
of Elasmobranchs, are simply embedded in the gums, and are only connected
with the jaws by fibrous tissue; but in some of the older fossil Sharks the
fixation of the teeth is effected by the mutual articulation of the basal
plates of the teeth with one another. The Chondrostean _Polyodon_, so
{250}shark-like in many other respects, also has teeth implanted basally in
the gums, and quite free from any special connexion with the jaw-bones. In
some Teleosts with movable teeth, the latter are merely attached to the
jaws by fibrous, and often elastic, ligaments, as in the Pike (_Esox_) and
the Angler-Fish (_Lophius_). As a rule, however, the teeth are directly
ankylosed to the bones developed in relation with the jaws. Very rarely,
as, for example, in some Characinidae, are the teeth implanted in sockets.
Nearly all Fishes are polyphyodont, that is, the old teeth are constantly
replaced by new teeth as fast as they become worn down or fall out. In the
Sharks and Dog-Fishes, for example, where the teeth are arranged in rows
parallel to the axis of each jaw, the functional teeth along the upper edge
of the jaw are usually erect, while those in the rows more internally
situated point inwards towards the oral cavity; and behind these again
there are rows of developing teeth in different stages of growth, and
partially hidden beneath a projecting fold of the oral mucous membrane
(Fig. 152). As the teeth in use become lost they are successively replaced
by the inner rows, which, with the mucous membrane in which they are
embedded, move forwards to the edge of the jaw, where they become erect and
functional. The teeth of the Holocephali and of the Dipnoi are not shed,
but the loss which they sustain through wear and tear is made good by
persistent growth at their bases. In the Teleostomi the succession is less
regular, new teeth being formed between or at the bases of the old teeth.
In the case of socketed teeth the succession is usually vertical, the new
teeth being formed at the sides of the old ones; and by the absorption of
the bases of the latter, the former come to lie directly below them, and
eventually they occupy the same sockets.
[Illustration: FIG. 152.—Transverse section through the jaw of a Shark
(_Carcharias_), showing how the teeth are replaced. _c_, Cartilage of the
jaw; _t_, functional tooth; _t′_, its immediate successor; _t"_, _t"_,
still younger teeth, covered by the fold of mucous membrane, _m. m_. (From
Ridewood.)]
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