The Cambridge natural history, Vol. 07 (of 10)Boulenger, George Albert
Science
The Cambridge natural history, Vol. 07 (of 10)
Boulenger, George Albert
Animals
There are also other considerations which add to the surplus power of the
extension stroke by lessening the resistance of the water to the flexion or
non-effective stroke. During the flexion stroke the tail fin is less
expanded and its area diminished, and by the rotation of the Fish on its
long axis the surface of the tail strikes the water obliquely, and further,
the tail moves with less rapidity. On the contrary, when the extension
stroke is made these conditions are reversed. The caudal fin is expanded,
the stroke is more rapid, and by the reverse rotation of the Fish the tail
now strikes the water with its flat surface. In other words, the action of
the tail during the two strokes may be compared to the "feathering" of an
oar in rowing. Nor is this all. A Fish in motion through the water produces
a suction current behind it. The current offers but little resistance to
the flexion stroke, inasmuch as the direction of the two coincide, but
during the extension stroke the tail meets the full force of the current,
and consequently its grip and propelling power are greatly enhanced. There
is a striking analogy between the movements of a Fish's tail in swimming
and the action of the screw of a steamer, but as a propelling organ the
former is far superior to the latter. As we have seen, the tail of a living
Fish can so adjust its shape and surface that it alternately eludes and
grips the water in accordance with the needs of particular strokes.
The curves into which the body of a Fish is thrown when swimming are never
less than two, but in long-bodied Fishes, such as the Eels, the number may
be increased, and in every case the curves occur in pairs and are
complementary to one another.
{353}Many Fishes can jump out of the water, either in pursuit of insect
food, like the Trout, or to enable them to escape the pursuit of their
foes, like the Flying-Fish (_Exocoetus_), by means of a single forcible
stroke of the tail, when the Fish is in a nearly vertical position close to
the surface of the water. It is thus that the Salmon executes its
remarkable leaps over weirs or up salmon-ladders when ascending rivers for
spawning.
The tail is also used for steering. If kept bent to one side when the Fish
is moving the tail acts like a rudder, and the course of the Fish is
deflected to that side; or the direction may be altered by single strokes
of the tail to the right or left, according to the course which the Fish
desires to pursue.
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