The Cambridge natural history, Vol. 07 (of 10)Boulenger, George Albert
Science
The Cambridge natural history, Vol. 07 (of 10)
Boulenger, George Albert
Animals
ANATOMY.—The more important points in the structure of the ascidiozooid of
_Pyrosoma_ are shown in Fig. 57. A circle of tentacles, of which one,
placed ventrally (_tn_), is larger than the rest, is found just inside the
circular branchial aperture. From this point a wide cavity, with a few
circularly placed muscle-bands running round its walls, leads back to the
large branchial sac (_br.s._), which occupies the greater part of the body.
The large stigmata are elongated transversely (dorso-ventrally), and are
crossed by internal longitudinal bars running antero-posteriorly. The
dorsal lamina is represented by a series of eight or ten {93}languets. The
nerve-ganglion (on which is placed a small pigmented sense-organ, the
unpaired "eye"), the neural gland, the dorsal tubercle, the peripharyngeal
bands and the endostyle are placed in the usual positions. On each side of
the anterior end of the branchial sac, close to the peripharyngeal bands is
a mass of rounded mesodermal gland-cells (_l.o_), which are the source of
the phosphorescence. They are apparently modified leucocytes lying in
blood-sinuses. The alimentary canal is placed posteriorly to the branchial
sac, and the anus opens into a large peribranchial or atrial cavity, of
which only the median posterior part (_cl_), is shown in Fig. 57. The heart
(_Ht_) lies between the posterior end of the branchial sac and the
intestine, close to where the endostyle is prolonged outwards to form the
inner tube of the ventral stolon. The reproductive organs are developed
from a cord of germinal tissue which forms a part of every budding stolon,
and so establishes a continuity of origin between the ova of successive
generations of _Pyrosoma_. On the ventral edge of the body, immediately
behind the stolon, with part of which it is continuous, a portion of this
germinal tissue gives rise to a lobed testis (_tes_), and to a single ovum
surrounded by indifferent or follicle-cells.
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