The two North American genera _Chorophilus_ and _Acris_, and the Brazilian _Thoropa_, connect the Hylidae with the Cystignathidae, in so far as their finger-discs are very small, or even {187}absent, and their sacral diapophyses are only slightly dilated. On the other hand, it has to be emphasised that the possession of adhesive discs on the fingers and toes does not necessarily constitute a member of the Hylidae. That requires _the further combination of an arciferous sternum, with dilated sacral diapophyses and teeth in the upper jaw_. Finger-discs are easily developed, and still more easily lost. Those of the typical Hylidae are constructed as follows. The terminal phalanx is elongated, claw-shaped, swollen at its base. Between it and the penultimate phalanx lies an interphalangeal cartilaginous disc which projects ventrally below the end-phalanx, thus assisting the formation of the ventral pad, and the turning upwards of the whole disc-like phalanx like the claw of a cat. This peculiar motion can be well observed in Tree-frogs which are at rest upon a horizontal leaf, or, better still, upon a rough stone, when the creatures take good care to adjust their discs into a safe and easy position. The pad or disc itself is furnished with unstriped, smooth muscular fibres, the contraction of which produces one or more longitudinal furrows on the under side. When the disc is in action or adhering, being flattened to a smooth surface, the end-phalanx sinks into the cushion; when not in action, the cushion swells and the phalanx appears as a slight dorsal ridge. The disc is rich in lymph-spaces, and its surface contains mucilaginous glands.