The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
_Gordonia_ and _Geikia_, of the New Red Sandstone of Elgin, are known from
their skulls only, but these are so well preserved that there is no doubt
about their close relationship to the typical South African Dicynodontia.
The skull of _Gordonia_ is about 7 inches long and 4 inches high. The
canines (Fig. 54, D, p. 280) are reduced to short, but thick, conical
tusks. The most {311}remarkable feature is the very elongated
squamoso-jugal arch, which arises moreover from the dorsal end of the long
squamoso-quadrate pedicle. The two wide and long temporal fossae are
dorsally divided by narrow parietal crests. There is a distinct
interparietal bone, and the usual interparietal foramen. The choanae are
united and lie within the palatines, which themselves are united; the large
lateral palatal foramina are otherwise enclosed by the pterygoids,
quadrates, and laterally by the squamoso-jugal arch.
ORDER IV. PLACODONTIA.
These are the latest and last members of the Theromorpha, unfortunately
known from skulls only, from the Muschelkalk or Middle Trias of Germany and
Russia. The skull of _Placodus gigas_ is about one foot long, rather high
and triangular owing to the lateral expansion of the temporal arches, which
diverge posteriorly. The squamoso-jugal arch is very broad, and most of the
posterior border of the orbit is formed by the large postorbital bone. The
maxillary bone seems to extend back to beyond the level of the orbits. The
choanae lie behind the premaxillaries. The palatines and pterygoids are
fused in the middle line, forming a broad bony palate, which, owing to the
broad, posteriorly extended wings of the pterygoids, much resembles that of
the crocodiles. The teeth are very remarkable. There are two or three
stout, conical, or chisel-like teeth in each premaxillary bone, and three
to five broad and flat maxillary teeth; three pairs of huge, broad, and
quite flat teeth are crowded together and fill up the whole vomerine and
palatine portion of the palate. These crushing teeth indicate that
_Placodus_ probably lived upon hard-shelled molluscs, and this would be in
conformity with its occurrence in the Muschelkalk, which is a strictly
marine deposit and full of shells. Another closely allied genus is
_Cyamodus_, one species of which is known from Russia. The teeth are fewer
in number and not so large as those of _Placodus_.
{312}CHAPTER IX
CHELONIA–ATHECAE–THECOPHORA
_SUB-CLASS IV.–CHELONIA._
There is no mistaking a tortoise. The shell and the horn-covered toothless
jaws separate them from all other four-footed creatures.
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