The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
All Chelonians lay white EGGS, round or oval, according to their kind, but
the shape of the eggs of one set sometimes varies within the greatest
limits. The shell varies from a parchment-like, flexible, scarcely
calcareous cover to a hard, well-polished case. As a rule the eggs,
imbedded in the ground, are hatched after a few months, but in some of the
northern kinds, e.g. _Emys orbicularis_, the hatching is deferred until the
next spring, the embryo's development being arrested during the winter. How
such eggs, buried a few inches only below the surface, withstand the often
very severe North German and Russian winter is a mystery. Whilst the
plastron is generally flat, it is more or less concave in the males of many
species, notably in _Testudo_, _Cistudo_, and _Emys_.
The general conclusions which can be drawn from the present GEOGRAPHICAL
DISTRIBUTION of the Chelonia are as few and unsatisfactory as those
applying to the Crocodilia, since all the main groups of Chelonians, and
many more extinct families, occurred together in bygone ages in the same
countries, for instance in Europe.
{332}[Illustration: FIG. 70.–Geographical distribution of Cryptodirous
tortoises.]
[Illustration: FIG. 71.–Geographical distribution of Pleurodirous
tortoises.]
The marine forms are naturally cosmopolitan, but the _Testudinidae_ are
likewise cosmopolitan, except in the Australian region. The _Chelydridae_,
now restricted to North and Central America, occurred formerly also in
Europe. The _Pleurodira_, in Mesozoic times plentiful in Europe, India, and
North America, are now restricted to South America, Australia, and Africa;
the _Pelomedusidae_ to Africa, Madagascar, and South America; the
_Chelydidae_ to South America and Australia. In the latter country all the
Chelonians belong to the Chelydidae. The _Trionychoidea_, occurring since
the Cretaceous epoch in North America, in Early and Mid-Tertiary times in
Europe, are now restricted to North America, Asia, and Africa. The country
richest in Chelonians is America; North and Central America together
possessing representatives of all the families except the Pleurodira, and
these we know to have died out there. The _Dermatemydidae_,
{333}_Cinosternidae_, and Chelydridae are now restricted to the Nearctic
sub-region (including Central America). Poorest in genera and species, all
of them Chelydidae, is the Australian region, where no fossils of other
families have yet been discovered. Europe, with its few Testudinidae, does
not come into consideration; Asia has at least Testudinidae and
Trionychidae, and in addition the solitary _Platysternum_ in Indo-China,
representative of a family whose affinities with the Chelydridae again
proclaim the validity of the Periarctic region.
[Illustration: FIG. 72.–Geographical distribution of Trionychidae and
Chelydidae.]
ORDER I. ATHECAE.
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