The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
The skull superficially resembles that of _Chelone_, chiefly owing to the
completely roofed-in temporal region. The supraoccipital crest is rather
short, covered completely by the parietals, the posterior margin of which
is rounded off instead of forming, as in the Chelonidae, a long projecting
triangular crest with the supra-occipital. The parietals are in broad
contact with the postfrontals, posteriorly they are just reached by the
squamosals. The quadrato-jugal is small, separated from the postfrontal by
the meeting of the squamosal with the jugal. The quadrate is notched
behind, and it separates the opisthotic from the squamosal. The
basisphenoid is large and broad, extending far forwards so as to separate
the pterygoids widely from each other except in their anterior portions,
which, instead of sending a lateral arm to the jugal and maxillary, as in
_Chelone_, are widely separated from these bones by the palatines. The
choanae lie on either side of the anterior half of the vomer, and are not
roofed over by ventral vomero-palatine wings.
The limbs and their girdles are essentially like those of the Chelonidae,
but are not derivable from them. The most remarkable feature is the shell.
The dorsal and ventral halves are directly continuous, forming one unbroken
case all round, which is composed of many hundreds of little bony plates,
irregularly polygonal, fitting closely into each other with their sutural
edges, and giving the shell a beautiful mosaic appearance. On the dorsal
side are a median row and three pairs of lateral rows of larger plates, and
these form seven longitudinal blunt ridges which all converge towards the
triangularly pointed tail-end of the shell. The ridges are not so much
produced by thickened {336}or spine-like edges of the plates, but by the
right and left halves of the plates being actually bent at an angle. This
is most conspicuous at the sides of the shell where it passes into the
ventral portion. The latter has two pairs of lateral and one median ridge.
The whole shell has consequently twelve ridges. The mosaic plates are
deeply imbedded in the cutis, being externally as well as internally
covered or lined with dense leathery skin. The epiderm is thin, and shows
no indications of horny scales. In young specimens the whole shell is soft
and very imperfectly ossified, later on it is quite rigid, although
comparatively thin. It is nowhere in contact with the internal skeleton,
except by a nuchal bone, which by a descending process articulates with the
neural arch of the eighth cervical vertebra.
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