The Cambridge natural history, Vol. 08 (of 10)Gadow, Hans
Science
The Cambridge natural history, Vol. 08 (of 10)
Gadow, Hans
Animals
_Dissorophus multicinctus_, also from the Permian of Texas, has been
described by Cope[40] as a "Batrachian Armadillo," and considered allied to
_Trimerorhachis_. Ten vertebrae are known, of an aggregate length of 93
mm.; the length of the creature was perhaps one yard. The neural spines are
elevated, and the apex of each extends in an arch on each side to the ribs.
These spinous branches touch each other, forming a carapace. Above, and
corresponding to each of them, is a similar dermal and osseous element,
which extends from side to side without interruption in the median line,
forming a dermal layer of transverse bands which correspond to the skeletal
carapace beneath it. This creature remotely approaches the genus
_Zatachys_, Cope, where a dermosteous scute is co-ossified with the apex of
the neural spine. The systematic position of this genus is at least
doubtful.
_Archegosaurus decheni_ from the Lower Red of Germany, known by many
well-preserved specimens, reached a length of 4 or 5 feet. The trunk
vertebrae are tri-partite, those of the tail quadri-partite, like those of
the trunk of _Chelydosaurus_. Young specimens show traces of gill-arches.
The thoroughly terrestrial walking limbs have four fingers and four toes;
the arrangement of the tarsalia, most of which are ossified, lend support
to the view that the morphological axis went through femur, fibula,
intermedium, the centralia, the second distal tarsale, and the second toe.
The dentine and enamel of the teeth are much folded, and this feature,
which applies to most members of this Order, to a lesser degree also to
others, has caused them to be comprised under the name of LABYRINTHODONTA.
The upper {83}surface of the head shows very characteristically arranged
grooves, which probably contained slime-canals and possibly sensory organs.
_Actinodon_ and _Euchirosaurus_ are closely allied forms, chiefly from the
Lower Red Sandstone of France; _Gondwanosaurus_ occurs in the Permian of
India.
ORDER III. STEGOCEPHALI STEREOSPONDYLI.
These are the most highly developed members of the typical Labyrinthodonta,
characterised by their much-folded teeth, and by their solid, bi-concave
vertebrae. _Loxomma_ occurs in the Upper Carboniferous of England and in
the Lower Red of Bohemia: _Trematosaurus_, _Capitosaurus_, and _Metopias_
from the New Red or Lower Trias to the Keuper of Germany. _Mastodonsaurus_
from the Trias of England and Germany is the most gigantic Amphibian known,
with a skull of nearly 1 yard in length.
_Labyrinthodon_ from the Keuper of Warwickshire is one of the latest
members of the group. Labyrinthodont creatures have also been described
from the Trias of South Africa, _e.g._ _Rhytidosteus_; those from North
America are insufficiently preserved.
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