The Cambridge natural history, Vol. 09 (of 10)Evans, A. H. (Arthur Humble)
Science
The Cambridge natural history, Vol. 09 (of 10)
Evans, A. H. (Arthur Humble)
Animals; Birds
Fam. I. RALLIDAE.–The Rails constitute a somewhat generalized and very
homogeneous Family, found in almost all parts of the world. The body is
peculiarly compressed–enabling them to move with ease in dense
vegetation–while the keel of the sternum is especially reduced in those
flightless forms for which the group is remarkable. The strong bill varies
in dimensions, being long in typical Rails, shorter and thicker in Crakes,
decidedly curved in _Himantornis_, and reaching its maximum size among the
Gallinules in _Porphyrio_ and _Notornis_, where it is subconical. A horny
shield is present upon the forehead in _Megacrex_, _Habroptila_, the
Gallinules and the Coots, which is usually rounded or truncated
{244}posteriorly, but is reduced to a point in _Porphyriops_. This
excrescence is in most cases red, but is sky-blue, light green, or dusky in
_Porphyriola_, green in _Tribonyx_, blackish in _Megacrex_, white, yellow,
or brown in _Fulica_. The lower part of the tibia is bare; the anteriorly
scutellated metatarsus is seldom short, though occasionally very stout; the
toes are long and slender with the elevated hallux weakest; the claws are
fairly long, curved, and sharp. Somewhat shorter digits are found in
_Tribonyx_ and _Pareudiastes_, _Fulica_ has broad lobes of skin along the
front toes, while _Porphyriops_ and _Gallinula_ have narrow entire
membranous margins to them. The wings are generally short and rounded, with
ten or eleven primaries, and from eleven to sixteen secondaries, all the
feathers being obtuse; but in many species these members are imperfectly
developed, and their coverts actually hide the quills in such cases as
_Ocydromus_ and _Notornis_. This retrograde tendency is clearly evidenced
in the "Island Hen" of Tristan da Cunha (_Gallinula_ or _Porphyriornis
nesiotis_) and the Mountain Cock of Gough Island (_G. comeri_), which
flutter along without flying; in the Moho of Hawaii (_Pennula ecaudata_),
_Ocydromus_ and _Notornis_ of New Zealand, and _Habroptila wallacii_ of
Halmahera; not to mention _Eulabeornis_, _Porzanula_, _Nesolimnas_,
_Cabalus_, _Pareudiastes_, and the extinct _Aphanapteryx_, _Aptornis_,
_Diaphorapteryx_, and _Erythromachus_. In several flightless forms, as in
the Dodo, the angle between the scapula and the coracoid is obtuse. The
tail has from ten to fourteen rectrices, the usual number being twelve;
these are short and usually soft, frequently with decomposed webs, and may
be concealed by the coverts, as in _Megacrex_, _Amurolimnas_, and
_Pennula_. Its form varies from narrow and pointed to comparatively broad
and rounded. A large caruncle rises behind the frontal shield in
_Gallicrex_ and _Fulica cornuta_, two knobs being found there in _F.
cristata_: the wing, moreover, is often armed with a sharp spine. The nasal
grooves are commonly long and deep; the pervious nostrils being in the hard
sheath of the bill in Gallinules, and partially covered by a bony or horny
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