The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
The chief facts which argue a primitive position among mammals for the
Cetacea are perhaps: (1) the slight union of the rami of the lower jaw; (2)
the occasionally rather marked traces of the double constitution of the
sternum; (3) the long and simple lungs; (4) the retention of the testes
within the body-cavity; (5) the occasional presence (in _Balaenoptera_) of
a separate supra-angular bone. These points, however, are but few, and are
not of such great weight as those which ought to be present to establish a
claim to separate treatment for the Cetacea as opposed to the Eutheria. If
this group of mammals can be tacked on anywhere, it appears to us that the
nearest relatives are not, as is sometimes put forward, the Ungulata or the
Carnivora, but the Edentata. There are quite a number of rather striking
features in which a likeness is shown between these apparently diverse
orders of mammals. The chief ones are these: (1) the existence of traces of
a hard exoskeleton, of which vestiges remain in the Porpoise; (2) the
double articulation of the rib of the Balaenopterids to the sternum, with
which compare the conditions obtaining in the Great Anteater; (3) the
concrescence of some of the cervical vertebrae; (4) the share which the
pterygoids may take in the formation of the hard palate; (5) the fact that
in the Porpoise, at any rate, as in many Edentates, the vena cava, instead
of increasing in size as it approaches the liver, diminishes.
Another group which is perfectly isolated is that of the Sirenia. The
alliance advocated by some with the Cetacea, and quite recently renewed by
Professor Haeckel, is contradicted by so many important features that it
seems necessary to abandon it. The recent discovery of a fossil Sirenian
jaw by Dr. Lydekker with teeth highly suggestive of those of Artiodactyla,
may prove a clue. A third group which is so isolated as to have been placed
in a {121} primary division, proposed to be called Paratheria, is that of
the Edentates. Probably the group so called should really be divided into
the Edentata and the Effodientia, the latter containing the Old World
forms. Whether or not it be ultimately shown that the Ganodonta are
ancestral Edentates (_sensu strictiori_), the connexion of the group with
others is not at present plain. The same is the case with the extensive
order of Rodents. It is true that the extinct order of the Tillodontia
shows certain Rodent-like characters on the one hand, and likenesses to
Ungulates on the other. Certain likenesses shown by such apparently diverse
animals as the Rabbit and the Elephant used to be insisted upon by
Professor Huxley. For the present, however, the Rodents must remain as an
isolated group with only very dubious affinities to others. The remaining
groups of existing mammals are easier to connect. At first the differences
between a Cat and a Horse seem to be quite as wide as those which separate
any two of the higher Eutherian orders. But it seems to become clearer and
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