The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
Of the Scaly Anteaters, Group SQUAMATA or MANIDAE, there is really but one
genus, though _Phatagin_, _Pholidotus_, _Smutsia_, and _Pangolin_ have been
used to distinguish various forms. The genus _Manis_ is African and
Oriental in range. Dr. Jentink, who has lately revised the species, allows
seven.[117] The external form of these animals is fairly well known, the
remarkable scales distinguishing the Pangolins from other animals. Between
the scales lie hairs, which seem to be absent in the adults of the African
species, though present in the young, thus affording a convenient method of
distinguishing the Ethiopian from the Oriental forms. The scales have been
compared to agglutinated hairs. That they are not "merely mimetic of the
Lizards' scales" is held by Weber,[118] who compares them directly with
those {189} structures, as he does the scales of other mammals, such as
those upon the tail of _Anomalurus_, etc. This, however, is not a universal
opinion. It is true that these scales occur chiefly in the lower forms of
mammals such as those under consideration, Marsupials, Rodents, and
Insectivores; but the fact that the hairs are developed before the scales
shows, or seems to show, that the former are the older structures, and to
lead to the inference that the scales of mammals are new structures. The
scattered hairs of the Pangolin have no sebaceous glands excepting on the
snout. This, again, looks as if they were degenerate structures, and
emphasises the non-archaic character of the scales. These animals have no
trace of teeth except possibly some slight epithelial thickenings which
have been interpreted as a last remnant; the tongue is suited for the
capture of ants, and is therefore much like that of the not nearly-related
American Anteaters. The stomach is of simple form; it is characterised by a
large gland, which suggests that of the Koala (see p. 144); the intestine
has no caecum. Retia mirabilia occur on the limb arteries. The placenta is
non-deciduate and diffuse; it is specially compared by Weber with that of
the Horse. Considering the many adaptive resemblances between this genus
and the American Anteaters, especially in the mouth cavity, it is
remarkable that in _Manis_ the pterygoids are not joined as they are in
_Myrmecophaga_. In spite of statements to the contrary, it appears that
there is sometimes a distinct lachrymal.
A remarkable feature in the skeleton of _Manis_ is the singular sternum.
The xiphoid cartilage is extraordinarily elongated into thin strips, which
reach the pelvis and return. This state of affairs is to be found in the
African species only. This structure is not comparable, as it has been said
to be, with abdominal ribs such as those of the reptile _Hatteria_.
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