The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
The curious form of teeth known as "selenodont" is characteristic of the
Artiodactyla, though only found well developed in the modern forms, and of
those only in the Pecora. The more primitive forms had "bunodont" teeth
with typically four tubercles (if we except the tritubercular and but
little-known _Pantolestes_); and the intermediate "buno-selenodont" type
characterises such groups as the Anthracotheriidae.
While the stomach of the Perissodactyles is always a simple sac, it is
complicated, or shows signs of complication, in the Artiodactyles. That of
the Hippopotamus is divided into two chambers; there are three in
_Tragulus_, and four in the typical Ruminants such as _Cervus_, _Ovis_,
etc.
Had we to deal only with the still living genera of Artiodactyles, it would
be easy to sort them into two groups on the characters of the teeth; for
the Pigs and Hippopotamus are provided with tubercular molars; they are
bunodont. The Deer, Camels, Oxen, Giraffes, etc., have selenodont molars.
Besides, the latter are "Ruminants," and have a more complicated stomach.
The existing Chevrotains forbid a more trenchant division, since they are,
as will be pointed out in due course, somewhat intermediate in structure;
the feet are more Pig-like, and the stomach is not so typically Ruminant.
In any case such a division is prevented by certain extinct families which
are perhaps ancestral to both. They have teeth which are not quite bunodont
and not quite selenodont. These teeth have been termed buno-selenodont or
buno-lophodont.
The distribution of the living Artiodactyles presents us with some
interesting facts. The vast preponderance of species occurs in the Old
World--34 in America as against over 250 species {273} in Europe, Asia, and
Africa. The Neotropical region has no Oxen, or Sheep, or Antelopes. The
latter are confined to Africa, Asia, and certain parts of the Palaearctic
region; they are vastly more prevalent in Africa, where they take the place
of the totally absent Deer. The Pig tribe is almost entirely Oriental and
Ethiopian in distribution, only one form, the European Wild Boar, ranging
into the Palaearctic region; and the two species of Peccary are found in
both North and South America. Broadly speaking, the Ethiopian region is the
headquarters of the Artiodactyla. But the great island of Madagascar has
but one form of Artiodactyle, a Pig of the genus _Potamochoerus_.[182]
GROUP I.--_SUINA._
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