The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
On account of its complication the stomach[233] has been compared to that
of Ruminants--it has even been alleged that Whales "ruminate"--but the
comparison will not hold good. Nor, on the other hand, is there a very
close resemblance to the equally-complicated stomach of the Sirenians.
The Rorqual has a stomach with as few compartments as any. The only Whale
which appears to have fewer is _Balaena mysticetus_, where there are but
three. In the Rorqual the oesophagus opens into a more or less globular
sac; from the upper end of this, _i.e._ close to the entry of the
oesophagus, arises the second chamber, long and narrowish; then follows an
extremely short third sac, {348} then a larger fourth, after which comes
the dilated commencement of the small intestine. The latter might be
regarded as a chamber of the stomach were it not for the fact that the
ducts of the liver and the pancreas open into it. This represents one type
of the Cetacean stomach, which seems to be found in all Whales except the
Ziphioids. In the latter, the oesophagus opens into the first compartment
as usual; but the second division of the stomach arises not close to the
entrance of the oesophagus, but at the opposite end. It would seem,
therefore, as if the first division of the stomach, found in most Whales,
were missing in Ziphioids. This way of looking at the matter is confirmed
by the fact that in _Hyperoodon_ a remnant of the missing first stomach is
found in the shape of a small diverticulum of the oesophagus just before it
enters the stomach.
The essential difference between the Whale's and the Ruminant's stomach is
this: in the latter the stomach is primarily divided into two portions, of
which the first is non-digestive and is clothed with oesophageal
epithelium. The second, the abomasum, is the digestive region. The first
part is again divided into three compartments. In the Whales, on the other
hand, it is the digestive part which is again subdivided, while if the
first part is divided it is not markedly so as in the Ruminants.
Public-domain text, read in full here on John Shaqi.
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