The Cambridge natural history, Vol. 10 (of 10) — John Shaqi
The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
The brain case is small proportionately and rounded. The "face" is
therefore long, and in some cases, especially among the fossil forms of
Platanistidae, the rostrum is extraordinarily elongated. The asymmetry of
the Whale's skull is one of its most remarkable features; this, however, is
entirely limited to the Toothed Whales, and among them is more pronounced
in some forms than in others. Thus the Platanistidae and many Ziphioids are
not nearly so asymmetrical as the {350} Dolphins and, especially,
_Physeter_. This asymmetry affects particularly the premaxillae, the
maxillae, and the nasals. The base of the skull is symmetrical. The Whale's
skull has very long premaxillae which, however, do not, except in the
extinct Zeuglodonts, bear any teeth. The nasal bones, whether symmetrical
or the reverse, are very small in existing Whales, which arrangement,
together with the long and broad maxillary bones, removes the anterior
nostrils, the blow-hole, far backwards. The roof of the skull is not at all
formed by the parietals externally. These bones form a portion of the side
of the cranium, but are replaced or covered by the {351}
enormously-developed supra-occipital in the adult. Here again the
Zeuglodonts are more typically Mammalian, for in them the parietals have a
normal development and situation, rising even into a median crest as in so
many quadrupeds. The bones related to the organ of hearing, the tympanis
and petrous bones, are very solid and dense in structure. Moreover they are
but loosely attached to surrounding bones, and are thus easily and
frequently lost. Nearly the only mammals which resemble the Whales in the
fact that the pterygoids sometimes meet in the middle line below are the
Edentata (Anteater and Armadillo, see p. 167). But in both groups this
peculiarity is not universal.
[Illustration]
FIG. 185.--A section of a skull of a young Caa'ing Whale (_Globicephalus
melas_). x 1/5. _a_, Angle; _an_, anterior nares; _AS_, alisphenoid; _bh_,
basihyal; _BO_, basioccipital; _BS_, basispnenoid; _cd_, condyle; _cp_,
coronoid process; _ExO_, exoccipital; _Fr_, frontal; _id_, inferior dental
canal; _IP_, interparietal; _ME_, ossified portion of the mesethmoid; _Mx_,
maxilla; _Na_, nasal; _Pa_, parietal; _Per_, periotic; _Pl_, palatine;
_PMx_, premaxilla; _pn_, posterior nares; _PS_, presphenoid; _Pt_,
pterygoid; _s_, symphysis of mandible; _sh_, stylohyal; _SO_,
supra-occipital; _Sq_, squamosal; _th_, thyrohyal; _Vo_, vomer. (From
Flower's _Osteology_.)
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