The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
This family of Whales is again susceptible of division into the two
sub-families--Physeterinae or Sperm Whales and the Ziphiinae or Beaked
Whales. Professor P. J. van Beneden was strongly against any subdivision of
what is here regarded as a perfectly natural family, embracing the
Physeters and the Beaked Whales. There are, however, some reasons for the
subdivision. The Ziphiinae have a reduced series of teeth, never exceeding
two on each mandible, which contrasts with the fully-toothed mandibles of
both _Physeter_ and _Kogia_. The stomach of the Ziphioids is
extraordinarily complicated even for a Cetacean. The small head of the
latter group, which recalls in a curious way that of Mosasauroid reptiles
and some Dinosaurs, is in contrast to the {363} enormous head of the
Cachalot and the very fairly-developed skull of the "Pygmy Sperm Whale."
Both, however, furnish spermaceti, and in various osteological details come
near together. On the whole we incline towards separating the Cachalots
from the Ziphioids, and shall therefore commence with the former as being
in some respects the more primitive members of the family Physeteridae.
SUB-FAM. 1. PHYSETERINAE.--This sub-family may be thus defined:--Teeth in
lower jaw numerous. No distinct lachrymal bone. Stomach with only four
compartments (? as to _Kogia_).
Of this sub-family the best-known genus is _Physeter_, including the Sperm
Whale or Cachalot. Of other reputed species we shall speak later. The genus
is characterised in the first place by its large size--as much as 82 feet
of length have been assigned to _Physeter macrocephalus_; but Sir William
Flower thought that 55 or possibly 60 feet might be a better approximation
to the greatest length of the Cachalot. The head is enormous, a third of
the length of the body, and terminates in a massive and bluntish snout.
This is, however, not so abruptly truncated as is often represented in
figures. According to Messrs. Pouchet and Chaves,[246] it slopes forward
two metres beyond the end of the lower jaw; the mouth is thus ventral and
almost shark-like in position, as is the case also with the Pygmy Sperm
Whale, to be considered later. In connexion with this peculiar position of
the mouth, it has been asserted--Mr. F. T. Bullen figures it[247]--that the
Sperm Whale turns over upon its back to bite. The blow-hole is single, and
shaped like the sound-hole of a violin; it lies upon one side, and is not
median in position. The throat is grooved as in the Ziphioids by two
grooves. The dorsal fin is represented by a whole series of lowish humps,
decreasing in elevation from before backwards. The pectoral fins are not
large relatively speaking. The great square head is not occupied entirely
by the skull; the cavity lying above, which is of course traversed by the
tube ending in the blow-hole, is filled with the spermaceti, which is fluid
fat during the life of the animal. Spermaceti also occurs in other Whales;
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