The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
resemblance to the mammalian skull; in fact it explains the origin of two
condyles from the typical reptilian single condyle, and disposes of the
necessity for believing, with Huxley and others, the Amphibia to be on the
main line of mammalian evolution on account of their two condyles. The
general aspect of the skull in _Cynognathus_ has been {94} compared to that
"of _Thylacinus_ or _Dissacus_." No one can examine the actual sketches of
the skull of that Theriodont without endorsing that opinion. As a curious
detailed point of likeness to certain Mammalia may be mentioned "a small
descending process of the malar bone, which may be a diminutive
representative of the descending element of the malar seen in _Elotherium_,
_Nototherium_, _Diprotodon_, _Macropus_, certain Edentata, such as
_Glyptodon_, _Megatherium_, _Mylodon_, _Bradypus_, but unparalleled so far
as I am aware in fossil reptiles." (Osborn.) The zoologist cannot help
being impressed with the significance of small details of similarity, which
do not seem to be due in any way to surrounding conditions of life, and
thus referable to mere convergence, like the fish-like form of Whales and
Seals.
The rest of the skeleton of the Theriodontia is by no means so well known
as the skull and teeth. But from what is known, other mammalian characters
can be pointed out. Perhaps the most striking mammalian feature is to be
found in the scapula of _Cynognathus_. It is in this creature somewhat
narrow and elongated; but it has a well-marked spine, ending in a hooked
acromion. Now it is to be noted in support, so far, of the diphyletic
origin of mammals, that in the Monotreme, as in Whales indeed, the spine
forms the anterior border of the scapula, and is coincident with it, there
being thus no prescapula at all in the Monotreme, and only a trace of it in
certain Whales.[47] Whether the multituberculate _Tritylodon_ or
_Diademodon_ had a scapula after the Monotreme pattern is not known; but it
is clear that the scapula of the triconodont _Cynognathus_ is quite after
the pattern of the Eutherian scapula. Furthermore, Professor Seeley is of
opinion that the coracoid was relatively small, and indeed smaller than the
same bone in Edentates, and _a fortiori_ than in Monotremes. Another fact
of structure which points also, possibly, in the direction of a diphyletic
origin for the Mammalia, is the double-headed ribs of _Cynognathus_. As is
well known, the ribs of the Monotremata have only the central head, the
capitulum.
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