The Cambridge natural history, Vol. 10 (of 10)Beddard, Frank E. (Frank Evers)
Science
The Cambridge natural history, Vol. 10 (of 10)
Beddard, Frank E. (Frank Evers)
Animals; Mammals
THE MAMMALS OF THE CRETACEOUS PERIOD.--At one time there was a totally
inexplicable gap between the Jurassic and the basal Eocene, a series of
strata which occupy an enormous expanse of time in the history of the earth
having appeared to {101} be devoid of mammalian remains. This gap, however,
has been filled up by the discovery of mammalian remains in the North
American Laramic formation, which seems to be clearly of Cretaceous age.
Furthermore, it is held by some that the Purbeck beds are more properly to
be placed with the Cretaceous, which would then necessitate the
consideration under the present heading of some of the types already dealt
with; and if, as is suggested in the following section, the lowest
so-called Eocene beds are really referable to the Cretaceous, there is no
lack of mammalian remains in that period. And, moreover, it was in that
case the Cretaceous period which witnessed the evolution of the existing
orders of Placental mammals. Otherwise the mammalian remains of the
Cretaceous agree with those of the Jurassic. We find remains of the
Multituberculata in fragments of Plagiaulacidae and Polymastodontidae.
_Ptilodus_ is a genus which has two premolars; and _Meniscoessus_ is
another multituberculate from the same Laramic formation. The other
detached fragments of mammals are thought by Osborn to represent both
Placentals and Marsupials.
THE MAMMALS OF THE TERTIARY PERIOD.--Unless the lowest beds of the earliest
Tertiary period, the Eocene, such as the Torrejon of North America, should
be in reality referred to the Cretaceous, there is no evidence that the
modern groups of Mammalia existed before the present epoch of the earth's
history. It is probable, however, that the Eutheria as a group were
Mesozoic. The fossil jaws that have been considered in the last chapter may
quite probably be primitive Eutherians, or even divisible, as believed by
Professor Osborn, into Marsupials and Insectivores. In the Tertiary,
however, apart from the question as to the nature of the Puerco and
Torrejon formations, and as to certain South American strata whose fossil
contents have been investigated by Professor Ameghino, we find the first
traces of mammals definitely referable to existing orders, or to be
distinctly compared with existing orders. Since, however, representatives
of types which have obvious relationships to modern types appear in
considerable profusion in the very earliest strata of the Eocene, it seems
clear that much remains to be discovered in beds earlier than these.
Confining ourselves, however, to facts and to comparisons which can be made
on more than a few lower jaws and scattered teeth, which is practically all
that we {102} possess of earlier mammals, we must arrive at the general
conclusion that two of the existing larger groups of the Eutherian,
non-Marsupial, mammals were differentiated at quite the beginning of the
Eocene, and were represented by forms from which it is possible to derive
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