Among the hundred passengers who landed from the _Mayflower_ at Plymouth Rock, on the 22nd of December, 1620, was a God-fearing Quaker named John Howland. He seems to have been unmarried at the time of his emigration; or at any rate his wife, if he had one, did not accompany him on the expedition. He settled in the colony of Massachusetts Bay, and left behind him a numerous progeny, whose descendants are to be found at the present day in nearly every State of the Union. From him, we understand, the subject of this sketch claims descent. The father of Sir William was Mr. Jonathan Howland, a resident of Dutchess County, in the State of New York. The latter was in early life a farmer, but subsequently engaged in commercial pursuits at Greenbush, in Rensselaer County, on the west bank of the Hudson River. He died at Cape Vincent, Jefferson County, in the year 1842. The maiden name of Sir William's mother was Lydia Pearce. Her family resided in Dutchess County, and were well-known and influential citizens. This lady still survives, and has attained the great age of ninety-four years. Soon after the death of her husband she took up her abode in Toronto, where she has ever since resided. The subject of this sketch, who was the eldest son of his parents, was born at the town of Paulings, Dutchess County, New York, on the 29th of May, 1811. He was brought up to farm work, but early displayed an aptitude for commercial life. After attending at a public school, and afterwards for a short time at the Kinderhook Academy, he determined to embark in a mercantile career. In the autumn of the year 1830, when he was barely nineteen years of age, he came to Canada, and settled in the village of Cooksville, on Dundas Street, in the township of Toronto. Here he obtained a situation as assistant in a country store of the period. In this store was kept the post-office for the village, the management of which largely devolved upon his own shoulders. The postal system in this Province had not then been very elaborately systematized. The mails for the whole of the western part of the Province passed over this route. The mail-matter for the different offices was not classified, but thrown into a bag, from which each successive postmaster selected such matter as was addressed to his office. The state of the roads was generally such that the mails had to be carried on horseback. Young Mr. Howland's duties required him to get up at one o'clock in the morning to receive the mail, which arrived at Cooksville at that hour. He was accustomed to select the mail-matter himself from the bag, after which he would hand the outgoing mail to the carrier, who then passed on westwardly to Dundas and Hamilton. Such was the primitive method of handling His Majesty's mail in Upper Canada in the year of grace 1830. It is scarcely to be wondered at that Mr. Howland, after such practical experience of the necessity for reform, should have allied himself with