Sexual cytogenic reproduction is based upon gametes or mating germ
cells. Since complementary gametes are specialized for union with each
other to form a single synthetic cell, the zygote, the number of their
nuclear threads or chromosomes is reduced to one half (the _haploid
number_) at the time of maturation, so that the somatic or tissue cells
of the parent organism have double the number (the _diploid number_)
of chromosomes present in the reduced or mature gametes. Hence, when
the gametes unite to form a zygote, summation is prevented and the
diploid number of chromosomes characteristic of the given species of
plant or animal is simply restored by the process of syngamy or union.
The process by which the number of chromosomes is reduced in gametes is
called _meiosis_, and, among the metists, it is distinct from syngamy,
which, in their case, is a separate process called fertilization. Among
the protists, we have, besides fertilization, another type of syngamy
called conjugation, which combines meiosis with fertilization.
In sexual reproduction, we have three kinds of gametes, namely:
isogametes, anisogametes, and heterogametes. In the type of sexual
reproduction known as isogamy, the complementary gametes are exactly
alike both in size and shape. There is no division of labor between
them. Each of the fusing gametes is equally fitted for the double
function which they must perform, namely, the kinetic function, which
enables them to reach each other and unite by means of movement,
and the trophic function which consists in laying up a store of
food for the sustenance of the developing embryo. In anisogamy, the
complementary gametes are alike in shape, but unlike in size, and
here we have the beginning of that division of labor, upon which the
difference of gender or sex is based. The larger or female gamete is
called a macrogamete. It is specialized for the trophic rather than
the kinetic function, being rendered more inert by having a large
amount of yolk or nutrient material stored up within it. The smaller
or male gamete is called a microgamete. It is specialized for the
kinetic function, since it contains less yolk and is the more agile of
the two. In anisogamy, however, the division of labor is not complete,
because both functions are still retained by either gamete, albeit
in differing measure. In the heterogamy, the differentiation between
the male and female gametes is complete, and they differ from each
other in structure as well as size. The larger or female gamete has
no motor apparatus and retains only the trophic function. The kinetic
function is sacrificed to the task of storing up a food supply for the
embryo. Such a gamete is called a hypergamete or egg. The smaller or
male gamete is known, in this case, as a hypogamete or sperm. It has a
motor apparatus, but no stored-up nutrients, and has even sloughed off
most of its cytoplasm, in its exclusive specialization for the motor
function.
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