Even the _vermiform appendix_ of the cæcum, which since Darwin’s time
has served as a classic example of a rudimentary organ in man, is, in
reality, not a functionless organ. Darwin, however, was of opinion
that it was not only useless, but positively harmful. “With respect to
the alimentary canal,” he says, “I have met with an account of only a
single rudiment, namely, the vermiform appendage of the cæcum. ... Not
only is it useless, but it is sometimes the cause of death, of which
fact I have lately heard two instances. This is due to small hard
bodies, such as seeds, entering the passage and causing inflammation.”
(“Descent of Man,” 2nd ed., ch. I, pp. 39, 40.) The idea that seeds
cause appendicitis is, of course, an exploded superstition, the hard
bodies sometimes found in the appendix being fecal concretions and not
seeds—“The old idea,” says Dr. John B. Deaver, “that foreign bodies,
such as grape seeds, are the cause of the disease, has been disproved.”
(Encycl. Americana, vol. 2, p. 76.) What is more germane to the point
at issue, however, is that Darwin erred in denying the utility of the
vermiform appendix. For, although this organ does not discharge in man
the important function which its homologue discharges in grain-eating
birds and also in herbivorous mammals, it subserves the secondary
function of lubricating the intestines by means of a secretion from its
muciparous glands.
Darwin gives the _semilunar fold_ as another instance of a vestigial
organ, claiming that it is a persistent rudiment of a former third
eyelid or _membrana nictitans_, such as we find in birds. “The
nictitating membrane, or third eyelid,” he says, “with its accessory
muscles and other structures, is especially well developed in birds,
and is of much functional importance to them, as it can be rapidly
drawn across the whole eyeball. It is found in some reptiles and
amphibians, and in certain fishes as in sharks. It is fairly well
developed in the two lower divisions of the mammalian series, namely,
in the monotremata and marsupials, and in some higher mammals, as in
the walrus. But in man, the quadrumana, and most other mammals, it
exists, as is admitted by all anatomists, as a mere rudiment, called
the semilunar fold.” (_Op. cit._, ch. I, pp. 35, 36.) Here Darwin is
certainly wrong about his facts; for the so-called third eyelid is
not well developed in the two lower divisions of the mammalian series
(_i.e._ the monotremes and the marsupials) nor in any other mammalian
type. “With but few exceptions,” says Remy Perrier, “the third eyelid
is not so complete as among the birds; (in the mammals) it never
covers the entire eye. For the rest, it is not really perceptible
except in certain types, like the dog, the ruminants, and, still more
so, the horse. In the rest (of the mammals) it is less developed.”
(“Elements d’anatomie comparée,” Paris, 1893, p. 1137.) Moreover,
Darwin’s suggestion leaves us at sea as to the ancestor, from whom
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