_Propliopithecus_, or _Dryopithecus_? Hylobatic, or troglodytic
affinities? Such questions are scarcely the pivots on which the
world is turned! Nevertheless, we rejoice that Doctor Gregory has
again settled the former problem (provisorily, at least) to his own
satisfaction. More important, however, than that of the dentition of
_Dryopithecus_, is the crucial question of whether or not Palæontology
is able to furnish evidence of man’s genetic continuity with this
primitive pithecoid root. Certainly, no effort has been spared to
procure the much desired proofs of our reputed bestial ancestry. The
Tertiary deposits of Europe, Asia, Africa, America, and the oceanic
islands have been diligently ransacked for fossil facts that would
be susceptible to an evolutionary interpretation. The aprioristic
criterion that all large-brained men are recent, and all small-brained
men with recessive chins are necessarily ancient, has always been
employed in evaluating the fossil evidence. Notwithstanding all
endeavors, however, to bring about the consummation so devoutly
desired, the facts discovered not only fail to support the theory of
collateral descent, but actually militate against it. For assuming that
man and the anthropoid apes constitute two distinct lines of evolution
branching out from common Tertiary or pre-Tertiary stock, palæontology
should be able to show numerous intermediate fossil forms, not alone
for the lateral branch of the apes, but also, and especially, for the
lateral line connecting modern men with the common root of the primate
tree. But it is precisely in this latter respect that the fossil
evidence for collateral descent fails most egregiously. Palæontology
knows of many fossil genera and species of apes and lemurs, that might
conceivably represent links in a genetic chain connecting modern
monkeys with Tertiary stock, but it has yet to discover so much as a
single fossil species, much less a fossil genus, intermediate between
man, as we know him, and this alleged Tertiary ancestor common to apes
and men.
Not even catastrophism can be invoked to save this irremediable
situation; for any catastrophe that would have swept away the human
links would likewise have swept away the ape links. The presence of
many genera and species of fossil apes, in contrast to the absence
of any fossil genus or species of man distinct from _Homo sapiens_,
is irreconcilable with the theory of collateral descent. Such is the
dilemma proposed to the upholders of this theory by Wasmann, in the
10th chapter of his “Die Moderne Biologie” (3rd edition, 1906), a
dilemma, from which, as we shall see, their every attempt to extricate
themselves has failed most signally.
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