Whenever the appearance of a so-called annectant type is antedated by
that of the two forms, which it is supposed to connect, this fact is,
naturally, a deathblow to its claim of being the “common ancestor,”
even though, from a purely morphological standpoint, it should
possess all the requisites of an ancestral type. Commenting upon the
statement that a certain genus “is a truly annectant form uniting the
Melocrinidae and the Platycrinidae,” Bather takes exception as follows:
“The genus in question appeared, so far as we know, rather late in the
Lower Carboniferous, whereas both Platycrinidae and Melocrinidae were
already established in Middle Silurian time. How is it possible that
the far later form should unite these two ancient families? Even a
_mésalliance_ is inconceivable.” (_Science_, Sept. 17, 1920, p. 260.)
Certainty, therefore, with respect to the comparative antiquity of
the fossiliferous strata is the indispensable presupposition of any
palæontological argument attempting to show that there is a gradual
approximation of ancient, to modern, types. Yet, of all scientific
methods of reckoning, none is less calculated to inspire confidence,
none less safeguarded from the abuses of subjectivism and arbitrary
interpretation, than that by which the relative age of the sedimentary
rocks is determined!
In order to date the strata of any given series with reference to
one another, the palæontologist starts with the principle that, in
an undisturbed area, the deeper sediments have been deposited at an
earlier period than the overlying strata. Such a criterion, however,
is obviously restricted in its application to local areas, and is
available only at regions of outcrop, where a vertical section of the
strata is visibly exposed. To trace the physical continuity, however,
of the strata (if such continuity there be) from one continent to
another, or even across a single continent, is evidently out of
the question. Hence, to correlate the sedimentary rocks of a given
region with those of another region far distant from the former,
some criterion other than stratigraphy is required. To supply this
want, recourse has been had to _index fossils_, which have now
come into general use as age-markers and means of stratigraphical
correlation, where the criterion of _superposition_ is either absent
or inapplicable. Certain fossil types are assumed to be infallibly
indicative of certain stratigraphical horizons. In fact, when it
comes to a decision as to the priority or posteriority of a given
geological formation, index fossils constitute the court of last
appeal, and even the evidences of actual stratigraphical sequence and
of physical texture itself are always discounted and explained away,
whenever they chance to conflict with the presumption that certain
fossil forms are typical of certain geological periods. If, for
example, the superposed rock contains fossils alleged to be typical
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