A very great school of art of the Renaissance was that which took its rise in Southern Tuscany and the Romagna, of whom the three best-known representatives are Piero dei Franceschi, Luca Signorelli and Melozzo da Forli. Piero's influence on Perugino has already been spoken of. Berenson declares him "hardly inferior to Giotto and Masaccio in feeling for tactile values; in communicating values of force he is the rival of Donatello; he was perhaps the first to use effects of light for their direct tonic or subduing and soothing qualities; and, finally judged as an illustrator, it may be questioned whether another painter has ever presented a world more complete and convincing, has ever had an ideal more majestic, or ever endowed things with more heroic significance." Piero's two pupils, Melozzo and Signorelli, each of them starting, as Berenson says, with the heritage Piero left him, yet following the promptings of his own temperament and the guidance of his own genius, touched excellence in his own splendid way. Melozzo was the grander temperament, {66} Signorelli the subtler and deeper mind. Visitors to Loretto, who see the music-making angels in a cupola there, are likely to be surprised into an appreciation of the power of the painter to express something of the witchery of music. Berenson says of them: [Footnote 6] "Almost they are French Gothic in their witchery, and they listen to their own playing as if to charm out the most secret spirit of their instruments. And you can see what a sense Signorelli had for refined beauty, if, when seated with Guido's 'Aurora,' you will rest your eyes on a Madonna by him in the same pavilion of the Rospigliosi Palace." [Footnote 6: _Op. cit_, p. 81.]