The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and PlantsDawson, John William, Sir
Science
The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and Plants
Dawson, John William, Sir
Paleobotany; Paleontology
[Illustration: FIG. 107.—Devonian Lepidoganoid Fishes (_Diplacanthus_
and _Osteolepis_). After Page and Nicholson.]
[Illustration: FIG. 108.—Modern Dipnoi.
_a_, _Ceratodus Fosteri_. Australia. _b_, _Lepidosiren annectus_.
Africa.]
A new and interesting light has recently been cast upon some of the most
anomalous of the ancient fishes by the study of the now rare and
peculiar species of the group of Dipnoi. Two of these, belonging to the
genus _Lepidosiren_, are the “Mud-fishes” of the rivers of tropical
Africa and America (Fig. 108, _b._) These creatures have an elongated
and elegant form, and the body is covered with overlapping horny scales
like those of ordinary fishes; but the pectoral and ventral fins are
rod-like, and are supported by simple cartilaginous rays, while the
tailfin forms a fringe around the posterior part of the body. Unlike
ordinary fishes, they have lungs as well as gills, and their mouths are
armed with sharp, bony, beak-like teeth (Fig. 115), with which they can
inflict terrible bites on the small fishes and frogs which furnish them
with food. Their most remarkable habit is that of burying themselves in
the mud of dried-up ponds, thus forming a sort of water-chamber or
“cocoon,” in which they remain in a torpid state until the return of the
rainy season sets them free.
Another example of these Dipnoi is the Barramunda, or _Ceratodus_ of the
Australian rivers (Fig. 108_a_). This fish resembles the _Lepidosiren_ in
many essential points of structure; but its fins have lateral rays, and
are consequently of some breadth, though of peculiar form, and its mouth
is armed with flat, pavement-like teeth, wherewith it browses on aquatic
grasses.
[Illustration: FIG. 109.—Anterior part of the palate of _Dipterus_.
Showing the dental plates at _a_, Devonian.—After Traquair.]
These modern fishes have enabled us to understand several mysterious
forms met with in the older rocks. In the first place, they show the
meaning of certain flat-toothed fishes, like _Dipterus_ of the Devonian
(Fig. 109), _Conchodus_ of the Carboniferous (Fig. 110), and _Ceratodus_
of the Carboniferous and Trias (Figs. 111, 112), previously of very
doubtful character. These must all have been of similar structure and
habits with the Barramunda, which is thus the sole survivor, perhaps
itself verging on extinction, of a group of herbivorous fishes
introduced, it may be, contemporaneously with the first stream affording
the requisite vegetable food, and which have continued almost without
improvement or deterioration to the present time. These fishes are,
however, very closely connected with the Ganoids, and there are some of
these, with fringed fins and overlapping scales, which, while regarded
as true Ganoids, resemble the Dipnoi very closely.
[Illustration: FIG. 110.—Dental plate of _Conchodus plicatus_ (Dn.).
Coal-formation of Nova Scotia. _Acadian Geology_.]
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