The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and PlantsDawson, John William, Sir
Science
The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and Plants
Dawson, John William, Sir
Paleobotany; Paleontology
The Millepedes, like the Land-snails, were first found in the
Coal-formation of Nova Scotia, but species have since been discovered
not only in Illinois, but also in Great Britain and in Bohemia. In Nova
Scotia alone two genera and five distinct species have been found, all
in the interior of erect trees, to which these creatures probably
resorted for food and shelter (Fig. 126). All the species yet known are
allied to the modern Gallyworms, though presenting special features
which seem to separate them as a distinct family,[46] and were probably
vegetable-feeders. Some of the species have the peculiarity, unknown
among their modern successors, of being armed with long spines.[47] The
moist, equable climate and exuberant vegetation of the Coal-period would
naturally be very favourable to Millepedes, and it is likely that the
discoveries made as yet give but a faint idea of their actual abundance.
It is not improbable that they subsequently declined, as we know of none
between the Carboniferous and the Jurassic, and they do not seem to have
improved up to the modern period. The Carnivorous Myriapods, however, or
Centipedes proper, a higher and essentially distinct type, are not known
until much more recent times.
[Illustration: FIG. 126.—Millepedes. From the Coal-formation.
_a_, _Xylobius sigillariæ_ (Dawson). _b_, _Archiulus xylobioides_
(Scudder). Anterior segments. Enlarged, _c_, _X. farctus_ (Scudder).
Caudal portion. Enlarged.]
The insects of the Carboniferous as yet known, belong to three out of
the ten or more orders into which the class is divided. One of these is
represented by a number of species of Cockroach, another by May-flies
and a Dragon-fly, and another by some weevil-like Beetles. The Cockroach
is characterised by Huxley as one of the “oldest, least modified, and in
many ways most instructive forms of insects;” and both he and Rolleston
take its anatomy as typical of that of the class. That these creatures
should have abounded in the Coal-period we need not wonder, when we
consider the habits of those that infest our houses, and when we further
bear in mind the number of species, some of them two inches in length,
that exist in tropical climates. So many species of this family have
been found in the Coal-formation on both sides of the Atlantic,[48] that
we may fairly regard them as constituting one of its most characteristic
features, and as probably the oldest representatives of the order to
which they belong[49] (Fig. 127). There were also in the Coal-period
insects allied to the Locusts and to the Mantids, a carnivorous group.
One of the latter (_Lithomantis_), described by Woodward, is a
magnificent insect, not unlike some modern tropical species. It was
found in the Coal-formation of Scotland. A still larger species,
probably the largest insect known, has been described by Brongniart. The
May-flies (_Ephemeridæ_) are represented in the Carboniferous by several
very large species. That of which the wing is shown in Fig.
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