The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and PlantsDawson, John William, Sir
Science
The Chain of Life in Geological Time: A Sketch of the Origin and Succession of Animals and Plants
Dawson, John William, Sir
Paleobotany; Paleontology
Had we lived in the Carboniferous period, we might have supposed that
the line of the great Labyrinthodont Batrachians would have been
continued onward and elevated, perhaps, in the direction of the
Mammalia, to which some features of their structure point. But we should
have been mistaken in this. The Labyrinthodonts, it is true, extend into
the Trias; but there is perhaps a sign of their coming degradation in
the appearance in the Permian of the first known mud-eel, a humble
Batrachian form allied to the Newts and Water-lizards.[62] Their special
peculiarities are dropped in the Mesozoic in favour of those of certain
small and feeble lizard-like animals, appearing first in the
Carboniferous, and more manifestly in the Permian, and which are the
true forerunners, though they can scarcely be the ancestors, of the
magnificent reptilian species of the Mesozoic, which have caused this
period to be called “the age of reptiles.”
The leading reptilian animal from the European Permian has long been the
_Proterosaurus_, from the copper slates of Thuringia (Fig. 141), a
reptile of lizard-like form, with well-developed limbs, and attaining a
length of three or four feet. It resembles more nearly those large
modern lizards known as “Monitors,” than any other existing form. The
fore-limb represented in the figure foreshadows very closely the bones
of the human arm and hand. Besides this we find in the Permian certain
lizards (_Theriodonts_ of Owen) which present the remarkable and
advanced peculiarity already predicted by some Carboniferous
Microsauria,[63] of having distinct canine teeth, producing a division
into incisors, canines, and molars, in the manner of the Carnivorous
quadrupeds, which they seem also to have resembled in some other parts
of their skeletons. It is not impossible that the footprints in the
Permian sandstones of Scotland, which have been referred to tortoises,
were those of animals of this type. Cope has recently described from the
Permian of Texas a number of reptiles which have the complex dentition
of the Theriodonts, and others which simulate that of Herbivorous
mammals, by the possession of flat grinding teeth supposed to be adapted
to vegetable food.[64] The teeth of all these Permian reptiles were set
in sockets, also an advanced peculiarity. Thus already in the Permian,
before the final decadence of the Carboniferous flora, and while the
Palæozoic invertebrates still lingered in the sea, the age of reptiles
dawned, and gave promise of its future greatness by the assumption on
the part of reptilian species of structures now limited to the Mammalia.
[Illustration: FIG. 141.—Arm of _Proterosaurus Speneri_. Reduced.
Permian.]
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