The Chemistry of Plant LifeThatcher, Roscoe Wilfred
Science
The Chemistry of Plant Life
Thatcher, Roscoe Wilfred
Botanical chemistry
But while this unvarying habit of growth is one of the fixed laws of plant
life, there are occasional deviations from it. A plant which, under normal
conditions of growth, develops in a certain fixed way, when exposed to
unusual environmental conditions, may, and often does, alter its habit of
growth in what may metaphorically be said to be an attempt to adjust itself
to the new conditions. Numerous examples of this phenomenon might be cited.
Certain algæ, which grow normally in water at a temperature of 20° to 30°
and which are killed if the temperature rises above 45°, have been grown
for successive generations in water the temperature of which has been
gradually raised, until they produce apparently normal growth in water the
temperature of which is as high as 78°; also, certain types of algæ
normally grow in the water of hot springs at temperatures of 85° to 90°,
and others in arctic sea-water the temperature of which sometimes falls to
-1.8° and never rises above 0° C. This phenomenon of the adjustment of a
species of plants to new conditions, which in the case of farm crops is
sometimes called "acclimatization," is of common occurrence and is often
utilized to economic advantage in the introduction of new strains of crops
into new agricultural districts. Again, the normal development of plants
may be altered as the result of injury or mutilation. Thus, if the ear is
removed from the stalk of Indian corn, at any time after flowering, there
always results an abnormal storage of sucrose in the stalk, instead of the
normal storage of starch in the kernels. Similarly, midsummer pruning of
fruit trees generally results in the production of abnormally large number
of fruit buds on the remaining limbs. Many other familiar examples of
alteration of normal development in response to, or as the result of,
abnormal conditions of growth might be cited.
TYPES OF ADAPTATIONS
To designate these different alterations of normal growth, several
different terms have been used. Among these, "adaptation," "accommodation,"
and "adjustment" have been commonly used by different biologists. Sometimes
these are used interchangeably, and sometimes different terms are used to
designate different types of response to altered conditions of growth.
Inasmuch as there seems to be no generally accepted usage of these
different terms, only one of them, namely, the word "adaptation" will be
used here; and different manifestations of this phenomenon will be
distinguished by using appropriate adjectives, as "physiological
adaptations," "chromatic adaptations," "morphological adaptations," etc.
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