The Chemistry of Plant LifeThatcher, Roscoe Wilfred
Science
The Chemistry of Plant Life
Thatcher, Roscoe Wilfred
Botanical chemistry
In general, starch is the final product of photosynthesis in most green
plants; but there are many exceptions to this. The polysaccharides, which
are generally insoluble, must be broken down into the simpler soluble
sugars before they can be translocated to other organs of the plant for
immediate, or future, use. When they reach the storage organs, they may be
recondensed into insoluble polysaccharides, or stored as soluble sugars.
Examples of the latter type of storage are, sucrose in beet roots, glucose
in onion bulbs, etc. Sometimes, this habit of storage seems to be a species
characteristic; as potatoes store starch, while beets, growing in the same
soil and under exactly the same environment, store sugar. But in other
cases, the nature of the carbohydrate stored undoubtedly is correlated with
the external temperatures at the time of storage. It has been shown that
cold, which tends to physiological dryness, very frequently favors the
storage of sugars instead of starches. Thus, in temperate zones, among
aquatic, or moisture-loving plants, those species which hibernate during
the winter at the bottom of lakes or ponds and are killed by temperatures
below freezing, store starch and no sugar; while in the same ponds, the
species whose storage organs pass the winter above the level of the water
and can withstand temperatures as low as -7° C. contain sugar during the
winter months, even if they contain starch during warmer periods.
Similarly, sugars often appear in the leaves and stems of conifers during
the winter months, only to disappear, or be replaced by starch, when spring
approaches. This same phenomenon is noticeable in arctic plants, which
generally contain but small proportions of starch and relatively large
amounts of sugars.
Similarly, the phenomenon of the turning sweet of potatoes when exposed to
low temperatures has often been noted. The change of the starch in potato
tubers to sugar is most rapid at the temperature of 0° C., and ceases at
7°, or above. Also, if potatoes in which the maximum amount of sugar is
present (not over one-sixth of the total starch can be converted into
sugar) are exposed to a higher temperature the sugar soon disappears.
In general, however, it may be said that each particular species of plant
has its own particular preference for a specific carbohydrate as its
reserve food material, and elaborates the proper enzymes to make it
possible to utilize this particular carbohydrate for its metabolic needs.
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