The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
_Fritschia curtidentata_ Dawson. Above: Bones of skull and
anterior extremity, and bony rods of belly. Below: Bones of pelvis and
posterior extremity. Nearly natural size. Erect tree, Coal formation,
South Joggins, Nova Scotia. Photograph by Dawson, published through the
courtesy of Dr. Arthur Willey. Original specimen in the Peter Redpath
Museum of McGill University.]
If now we take up a consideration of each of the characters mentioned
by Lydekker we find that the first one holds good for all examples of
the group. The second character, "without limbs," is not good. Species
of _Œstocephalus_, _Ptyonius_, _Molgophis_ all possess limbs; and
doubtless Ophiderpeton will be found to possess limbs also, since it
has a well-developed pectoral girdle. The limbs in all of these genera
are small. The third character, "apparently without pectoral and pelvic
girdles," is not at all a good character, since nearly every specimen of
some species and almost all species show evidences of pectoral girdles
and a few exhibit pelvic girdles. The fourth character, "vertebræ with
elongated centra and aborted neural spines," is not a good distinguishing
character, since _Amphibamus_, an undoubted microsaurian, possesses
the same vertebral characters. "Ribs slender, and barbed like those of
fishes" is a character which is common to several widely distinct genera.
All Microsauria possess long, slender ribs, and the barbed condition
is one which is possessed by only a few, _Thyrsidium_, _Ophiderpeton_,
etc., the so-called "barb" being merely a highly exaggerated tuberculum.
The teeth of nearly all Microsauria are smooth, so that the character
"teeth smooth" is not a good one for a group definition. It has not been
possible to examine any of the American specimens for the plication of
the dentine, since the forms are so rare and the fossils very fragile.
The last character, "external gills probably persistent," is certainly
not true for the American species, and the evidence for the European
species is negative. Fritsch described and figured (Fauna der Gaskohle,
Bd. I, 1883, p. 114, Tafeln 18 and 23) structures which he regarded as
supporting structures for the external branchiæ. He says in regard to
these structures:
"Bei dem Umstande, dass sie von der Kiemengegend aus sich
büschelförmig verbreiten und man ihren Contakt mit einer Art von
Branchiæ constatiren kann, zweifle ich nicht daran, dass diese
Stäbchen dem Kiemenapparate angehören. Bedenklich ist nur ihre
grosse Zahl und das Vorkommen bis zum 16ten Wirbel und ich erwog
die Möglichkeit, dass diese Stäbchen einem zarten Bauchpanzer
angehören könnten. Da aber weiter im Verlaufe der ganzen Wirbelsäule
nichts Aehnliches vorkömmt, so ist man gezwungen anzunehmen, dass
_Dolichosoma_ sehr grosse lange Kiemenbüschel besessen haben muss."
Public-domain text, read in full here on John Shaqi.
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