The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
There are 5 species of Tuditanus thus far known. All of the species are
characterized by the possession of a peculiar triangular-shaped clavicle
with radiating grooves, and this has been taken as one of the distinctive
characters of the genus, as well as of the family. The structure of the
cranium where known is quite uniform among the different species. The
squamosal is quite large and the supratemporal is not always closely
joined to the parietal. The species are:
_Tuditanus punctulatus_ Cope, Linton, Ohio.
_brevirostris_ Cope, Linton, Ohio.
_longipes_ Cope, Linton, Ohio.
_minimus_ Moodie, Cannelton, Pennsylvania.
_walcotti_ Moodie, Linton, Ohio.
=Tuditanus punctulatus Cope.=
Cope, Trans. Amer. Phil. Soc., XV, p. 271, 1874.
Cope, Geol. Surv. Ohio, II, pt. II, p. 392, pl. xxxiv, fig. I, 1875.
Type: Specimen No. 110, American Museum of Natural History, where there is
also specimen No. 111.
Horizon and locality: Linton, Ohio, Coal Measures.
This species, together with the form, _T. brevirostris_, described on
p. 88, was used by Cope as the type of the genus _Tuditanus_. Cope
subsequently associated some reptilian remains from the Linton mines with
the type of _T. punctulatus_ and changed the generic term to _Isodectes_,
which was known by another species, _I. megalops_ Cope, from the Permian
of Texas. The remains associated by Cope with _I. megalops_ undoubtedly
represent a reptilian species and which has been described elsewhere
under the name _Eosauravus copei_ Williston. The species is of exceeding
interest because it is the _oldest known reptile_ and places the range of
the Reptilia down towards the base of the Pennsylvanian.
The species _Tuditanus punctulatus_ Cope was founded on well-preserved
remains of nearly the entire skeleton of a single individual. The bones
are represented by shining carbonaceous matter, and since both of the
slabs containing the impression were preserved, a great many characters
have been determined. The head, 1 fore-limb, and 23 consecutive vertebræ
with ribs are well defined, but of the pelvis and hind limbs nothing is
visible.
The cranium (fig. 19) is very similar to that of _T. minimus_, from
the Cannelton, Pennsylvania, slates. It is triangular in shape, with a
narrowed obtuse muzzle. The orbit of the left side is well defined and
lies well forward. It is oval in outline and its width is about two-thirds
of its length. The nostrils are small and are located well toward the tip
of the muzzle. The parietal foramen lies behind the median transverse line
which divides the skull equally.
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