The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
[Illustration: Fig. 21. A. Outline drawing of type of _Tuditanus walcotti_
Moodie, from the Coal Measures of Linton, Ohio, showing impression of body
and muscle at _M_. × 25. _cl_, clavicle; _fr_, frontal; _f_, femur; _h_,
humerus; _nos_, nostril; _or_, orbit; _par_, parietal; _rb_, rib; _pp_,
postparietal; _v_, vertebra; _pfo_, pineal foramen.
B. Left leg of second specimen of Tuditanus walcotti. × 3.]
The specimen includes, besides the body impression, the complete skull, a
right clavicle, with portions of the left, a left humerus, 12 cervical and
dorsal vertebræ, 10 pairs of ribs somewhat disturbed as to position, and a
portion of the mandible. There are no traces of ventral scutellæ nor body
scales in the smooth impression of the carbonized skin. One would expect
to find impressions of the ventral scutæ in this specimen if they were
present. Cope remarked on the apparent absence of scutellæ from members of
the genus _Tuditanus_ as they were known to him, and no contrary evidence
has since been brought to light. Until such evidence is forthcoming the
absence of scutes will be taken as one of the generic characters of the
genus _Tuditanus_. Under a magnification of 50 diameters the carbonized
skin shows as folds and wrinkles, like muscle fibers, in some places; in
others no traces of the muscular structure can be detected. The wrinkles
may be impressions of the internal musculature of the body-wall of the
abdomen. It is especially well preserved in the pelvic and pygal regions.
Sections of the coal were made, but nothing definite could be determined
as to the character of the impressions, as they were too poorly preserved
and the coal was too soft to bear much handling.
The specimen is preserved on the belly, with the dorsum of the skull
uppermost. It has been practically impossible to determine the arrangement
of any of the cranial elements except the f rentals, parietals, and
postparietals, which have the relations indicated in figure 21, A. A
median suture is clearly evident, with the pineal foramen well back in
this suture. The bones of the skull are marked with faint radiating lines.
It is in the form of the skull and the position of the orbits that the
specific characters are found, as follows: the backward position of the
eyes and the oval, pointed shape of the skull. The species is closely
related to _Tuditanus minimus_ Moodie, from the Cannelton slates of
Pennsylvania, and serves further to connect the forms from the Ohio and
Pennsylvania localities. It differs from the last-named species in the
position and form of the orbits, these structures being more oval in the
present form and placed further back. The shape of the skull differs also
in the almost entire absence of the posterior table. The median points of
the orbits occupy the line which bisects the skull, and the interorbital
width is less than the width of the orbit. The mandible is heavy and
appears to have borne sharp, pleurodont teeth.
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