The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
Zittel, Handbuch der Paleontologie, Bd. III, Abth. 1, p. 397, 1887.
Large terrestrial vertebrates; largest of the class. Skull equal to
one-fourth or one-third of the entire body in at least one species,
_Metoposaurus diagnosticus_ von Meyer (242). Lateral-line canals
always present (458) on the skulls as deeply impressed grooves which,
in life, were possibly roofed-over by a cartilaginous or other
connective-tissue membrane. The sensory organs undoubtedly being
supplied by the superficial ophthalmic branch of the trigeminal
nerve, branches of which pierced the cranial elements near the
grooves, no evidence of such openings in the bottoms of the grooves;
the condition probably being analogous to _Hydrolagus colei_ and
other chimæroids. Vertebræ stereospondylous, with well-developed
neural arches from which projected the well-developed zygapophyses,
sometimes slightly amphicœlous and pierced for the notochord, such
forms being uncertainly placed in the group. Tail unknown, possibly
short. Limbs and girdles well developed (243), phalangeal formula
unknown; carpus osseous and tarsus unknown. Pectoral girdle composed
of osseous scapulæ, clavicles, interclavicle, coracoid (?); clavicles
and interclavicle ventrally sculptured. Pelvic girdle composed of
osseous pubis, ischium, and ilium (242), the pubis a small plate, in
life largely cartilaginous, the three uniting by cartilaginous union
to form the acetabulum. Ventral armature unknown, possibly wanting.
Range: Pennsylvanian to Upper Triassic.
Distribution: North America, Europe, Asia, Africa, and Australia.
=Family MASTODONSAURIDÆ Huxley, 1863.=
Huxley, Quart. Jour. Geol. Soc., XIX, p. 65, 1863.
Lydekker, R., Cat. Fossil Reptilia Amphibia, pt. IV, p. 141, 1890.
Skull triangular, and more or less elongated, with the cranial bones
very strongly sculptured, the occipital condyles ossified (49), and
large palatal vacuities; dentine of teeth with very complex plications
(502); no bony rings (242) in sclerotic; and no ventral scutes.
Bodies of vertebræ (49) fully ossified in the adult. There are large
palato-vomerine tusks on the inner side of the maxillary teeth; and
the palatines run parallel to the maxilla. The mandible has a large
postarticular process; and there is a small inner series of mandibular
teeth. In the type genus the pubes are separate from the ischia, and
do not enter into the formation of the acetabulum; and the sacral ribs
form kidney-shaped disks (393).
Represented in North America by a single tooth from the Carboniferous of
Kansas. Described by Williston as Mastodonsaurus sp. (Kans. Univ. Quart.,
VI, pp. 209-210, pl. XXI). Represented in the Triassic of Wyoming by
Anaschisma browni, Branson (49).
=Mastodonsaurus sp. indet.=
Williston, Kans. Univ. Quart., VI, p. 209, 1897.
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