The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
Terminal phalanges sometimes clawed. Carpus and tarsus osseous or
cartilaginous, usually the latter. Ribs never attached to a sternal
apparatus, single or double headed or intermediate, long and curved
or short and straight. Articulation with vertebral column inter- or
intra-central. Respiration both branchial and pulmonary; branchiæ
persistent and osseous in some forms. Development by metamorphosis
either in the egg membrane, on the back of the mother, or in the
water. No amnion or allantois. Heart with a single ventricle and 3 or
4 pairs of aortic arches; postcava always present in the recent forms.
DEFINITION OF SUBCLASS EUAMPHIBIA, MOODIE, 1909. COAL MEASURES TO RECENT.
(World-wide distribution.)
Moodie, Trans. Kans. Acad. Sci., 1909, p. 243.
Moodie, Geol. Mag., n. s., Dec. V, vol. VI, p. 220, May, 1909.
The present group was established for the reception of the Branchiosauria
and their descendants, the Caudata, with the related forms, the Apoda. The
Salientia are included provisionally, since there is no evidence of the
origin or relationship of this group of animals to other Euamphibia save
that they have attained the same stage of evolution. They are in no way
closely related to any known group of Amphibia, recent or extinct, but
they stand on the same plane of development as the Caudata and present
similar structures, _i.e._, a single ventricle in the heart, external
branchiæ in the young, a glandular skin, perichondral bone, and a large
parasphenoid. The origin of the Salientia is a puzzle and must remain so
until further paleontological evidence is forthcoming. Wyman, Cope, and
the writer have all remarked on the similarity of structure between the
Salientia and the single known specimen of _Pelion lyelli_ Wyman from the
Linton, Ohio, Coal Measures.
The subclass Euamphibia may be defined as follows: Aquatic or terrestrial
Amphibia; development by metamorphosis; external branchiæ present in
the young; bones almost entirely perichondral; carpus and tarsus never
ossified; osseous pubis absent; vertebræ usually amphicœlous with
persistent notochord; ribs short and straight or flat and slightly curved,
or absent; digits 4 in the hand and 5 in the foot; skull never grooved or
pitted, nor cut by the lateral-line canals; lateral-line organs present in
the skin; sclerotic plates present or absent; tail long and flattened or
absent. Ribs in _Triton walthi_ are secondarily long and curved.
DEFINITION OF THE ORDER BRANCHIOSAURIA LYDEKKER, 1890. COAL MEASURES AND
PERMIAN OF NORTH AMERICA AND EUROPE.
Lydekker, Cat. Fossil Reptilia and Amphibia, pt. IV, p. 208, 1890.
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