The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
Fig. 16_a_. Nearly complete specimen of _Cocytinus gyrinoides_ Cope, from
the Coal Measures of Linton, Ohio. Original in the American Museum of
Natural History. × 0.95.
Measurements (No. 2564, American Museum of Natural History).
mm.
Length of entire specimen 113
Length of skull 15
Width of head posterior 15.5
Length of vertebra 2
Length of rib 4
=Genus ERIERPETON Moodie.=
Moodie, Kans. Univ. Sci. Bull., VI, No. 2, p. 328, 1912.
Type: _Erierpeton branchialis_ Moodie.
The generic characters are found, first of all, in the presence of
hyobranchial arches which indicate its relationship to the formerly
described _Cocytinus gyrinoides_ Cope, from Ohio. The only other known
extinct genera of Caudata which possess, or at least have preserved, the
hyobranchial arches are the Jurassic _Hylæobatrachus_ from Belgium and
Lysorophus from the Permian of Texas. The present form is widely distinct
from both of these genera in the shape of the mandible and the form and
arrangement of the hyobranchial bars. The genus _Erierpeton_ finds its
closest ally in _Cocytinus_, in the family Cocytinidæ, which possibly
belongs in the order Caudata and the suborder Proteida of Cope.
=Erierpeton branchialis Moodie.=
Moodie, Kans. Univ. Sci. Bull., VI, No. 2, pp. 329-330, pl. 1, fig. 3;
pl. 2, fig. I, 1912.
Type: Specimen No. 801 (222) 5, Yale University, Museum.
Horizon and locality: Mazon Creek shales, near Morris, Illinois.
The amphibian remains designated by the above name consist of a distinct
mandible and some rather indefinite body impressions (plate 3, fig.
4). Three elongate impressions occur between the rami of the mandibles
(fig. 15, A), which, I suppose, must represent hyoid bones belonging to
the hyobranchial arches. The lateral elements are paired and the median
impression is straight and lies between the paired impressions of the
hyoids. The paired portions probably represent the hypohyals or hypohyals
plus the ceratohyals, and the unpaired portion of the first basibranchial,
according to the nomenclature of Wiedersheim (Comparative Anatomy of
Vertebrates, 1897, p. 86). If the impressions have been correctly
interpreted the present specimen is of very great interest, since it is
the first evidence we have of the hyobranchial arches in the Amphibia
of Mazon Creek, and the second in the Carboniferous of North America.
Dawson doubtfully identified (216) some elements of the Joggins Amphibia
as hyoids, but was uncertain as to their position. Cope described fully
the well-developed hyobranchial apparatus of _Cocytinus gyrinoides_ (123)
from the Coal Measures of Ohio. Among other Paleozoic Amphibia Williston
(614) has described branchial arches in the peculiar form _Lysorophus
tricarinatus_ Cope, from the Permian of Texas.
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