The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
The order is suggested in the Coal Measures by a single species, known
from a single poorly preserved specimen (plate 24, fig. 1). The form
_Pelion lyelli_ Wyman was the first known of the Linton Amphibia, and
its striking frog-like (123, 639) appearance was early noticed. There
is no assurance that the species belongs with this order, but since a
well-developed and highly specialized frog (480, 481) occurs in the Como
Beds (405) of Wyoming, it is not impossible that we may have in the
_Pelion lyelli_ a suggestion (460), at least, of the ancestral structure.
It is certain that the frogs have, in past ages, had a much greater length
of vertebral column than they possess at present, as is witnessed by the
coalescence of several vertebræ to form the urostyle. It is suggested that
the ancestral vertebral column is represented in _Pelion_.
=Family PELIONTIDÆ Cope, 1875.=
Cope, Bull. U. S. Nat. Mus., No. 1, p. II, 1875.
The present family includes but a single species, that of _Pelion lyelli_
Wyman, first described in 1858 (640), from Linton, Ohio.
The family characters are to be found in the broad and obtusely rounded
cranium, in the frog-like scapular arch, the frog-like hind limb, and in
the form of the palate, so far as these structures have been preserved.
It has been suggested that the present form shows decided affinities with
the frogs of to-day and it may possibly be looked upon as the actual
ancestor of the living frogs. The length of the vertebral column would
seem to militate against such a relationship, since it is well known
that frogs have had a short vertebral column since the Jurassic (480,
481). But this is not a good argument, since the developing urostyles of
modern tadpoles show metameric fenestrations in the developing bone which
doubtless correspond to openings between the vertebræ. The notochord
of the tail is segmented, apparently through the influence of former
vertebral structure. At any rate, the suggestion is an interesting one
and, whether sustained or disproven, the present discussion is based on
the probabilities of the case.
=Genus PELION Wyman, 1868.=
Wyman, Am. Jour. Sci. (2), XXV, p. 160, 1858 (_Raniceps_).
Cope, Proc. Acad. Nat. Sci. Phil., 1868, 211 (_Pelion_, suggested in
letter to Cope by Wyman).
Type: _Pelion lyelli_ Wyman.
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