The Coal Measures Amphibia of North AmericaMoodie, Roy Lee
Science
The Coal Measures Amphibia of North America
Moodie, Roy Lee
Amphibians, Fossil -- North America; Paleontology -- Carboniferous; Paleontology -- North America
This is the most frog-like, in appearance at least, of all the Amphibia
which have so far been discovered in the Carboniferous. The skull
especially has a shape which is strikingly frog-like, and the long hind
limbs lend further likeness to the tailless forms. _Pelion_ may have
been a jumping creature, if we may judge from its long hind legs. Wyman
and Cope have both called attention to the frog-like appearance of the
specimen, and this is apparent at the first glance. It is probable that
the resemblance has some significance as to the ancestry of the Salientia,
and it may indicate the first step in the origin of the tailless Amphibia.
It is possible that the frogs began to be separated from the other
Amphibia during the Carboniferous. The first frogs we know are from the
Jurassic, where they are well-developed ranids. If _Pelion_ be a frog
ancestor, then the history of the group from the Coal Measures to Jurassic
is an unknown story.
The specimen is preserved on its back and it is thus impossible to tell as
to the structure of the skull. Cope was of the opinion that the depressed
areas on the sides of the elongate parasphenoid were the orbits, and if
so the resemblance to the frogs is much more striking. In the frogs there
is a strong process from the pterygoid which projects inward to meet a
corresponding process from the parasphenoid. This forms a heavy rod behind
the palatine vacuity. There is a heavy rod represented in the specimen and
a part of it is certainly the external process of the parasphenoid, but
whether it is to be interpreted as in the frog is an open question. The
outline of the cranium is partially obscured by the mandibles, but the
anterior part is represented by a raised line, as shown in figure 17. In
the anterior part of this space there are two ridges which may be tooth
ridges. If they are teeth there is a great similarity to the premaxillary
and vomerine teeth of _Necturus_, since the ridges are widely separated
at the median line and approximated distally, as they are in _Necturus_.
The mandible is preserved entire and its form is strikingly frog-like. Its
posterior angles project over the quadrate area and seem to have had an
upturned projection such as is found in the mandibles of the Crocodilia.
There are impressions of 20 vertebræ preserved, and they cover a little
more than half of the presacral region. There may have been 28 to 30
presacrals. The vertebræ, as preserved, are somewhat quadrate in outline
and constricted at the middle, as though they were of the typical
microsaurian type. No ribs are preserved.
[Illustration: Fig. 17. _Pelion lyelli_ Wyman, an amphibian from the
Coal Measures of Ohio, the supposed ancestral salientian. (After Wyman.)
× 0.75. _pmx_, premaxilla; _pv_, palatine vacuity; _m_, mandible; _sc_,
scapula-coracoid; _h_, humerus; _r-u_, radius and ulna; _f_, femur; _t_,
tibia.]
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