The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
Immediately beneath the epiostracum, there is a zone {194} which
may occupy a sixth or a seventh of the thickness of the whole, which
is more transparent than the rest, and often presents hardly any
trace of horizontal or vertical striation. When it appears laminated,
the strata are very thin. This zone may be distinguished as the
_ectostracum_ (_b_), from the _endostracum_ (_c_), which makes up the
rest of the exoskeleton. In the outer part of the endostracum, the
strata are distinct, and may be as much as 1‐500th of an inch thick,
but in the inner part they become very thin, and the lines which
separate them may be not more than 1‐8000th of an inch apart. Fine,
parallel, close-set, vertical striæ (_e_) traverse all the strata of
the endostracum, and may usually be traced through the ectostracum,
though they are always faint, and sometimes hardly discernible, in
this region. When a high magnifying power is employed, it is seen
that these striæ, which are about 1‐7000th of an inch apart, are
not straight, but that they present regular short undulations, the
alternate convexities and concavities of which correspond with the
light and the dark bands respectively.
If the hard exoskeleton has been allowed to become partially or
wholly dry before the section is made, the latter will look white
by reflected and black by transmitted light, in consequence of the
places of the striæ being taken by threads of air of such extreme
tenuity, that they may measure not more than 1‐30,000th of an inch in
diameter. It is to be concluded, therefore, that {195} the striæ are
the optical indications of parallel undulating canals which traverse
the successive strata of the cuticle, and are ordinarily occupied by
a fluid. When this dries up, the surrounding air enters, and more or
less completely fills the tubes. And that this is really the case
may be proved by making very thin sections parallel with the face of
the exoskeleton, for these exhibit innumerable minute perforations,
set at regular distances from one another, which correspond with the
intervals between the striæ in the vertical section; and sometimes
the contours of the areæ which separate the apertures are so well
defined as to suggest a pavement of minute angular blocks, the
corners of which do not quite meet.
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