The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
The rostrum grows out between the procephalic lobes; it remains
relatively very short up to the time that the young crayfish quits
the egg, and is directed more downwards than forwards. The lateral
portions of the carapacial ridge, becoming deeper, are converted into
the branchiostegites, and the cavities which they overarch are the
branchial chambers. The transverse portion of {218} the ridge, on
the other hand, remains relatively short, and constitutes the free
posterior margin of the carapace.
As these changes take place, the abdomen and the sternal region of
the thorax are constantly enlarging in proportion to the rest of
the ovum; and the food-yelk which lies in the cephalothorax is,
_pari passu_, being diminished. Hence the cephalothorax constantly
becomes relatively smaller and the tergal aspect of the carapace less
spherical; although, even when the young crayfish is ready to be
hatched, the difference between it and the adult in the form of the
cephalothoracic region, and in the size of the latter relatively to
the abdomen, is very marked.
The simple bud-like outgrowths of the somites, in which all the
appendages take their origin, are rapidly metamorphosed. The
eyestalks (fig. 59, _1_) soon attain a considerable relative size.
The extremities of the antennules (_2_) and of the antennæ (_3_)
become bifurcated; and the two divisions of the antennule remain
broad, thick, and of nearly the same size up to birth. On the other
hand, the inner or endopoditic division of the antenna becomes
immensely lengthened, and at the same time annulated, while the outer
or exopoditic division remains relatively short, and acquires its
characteristic scale-like form.
The labrum (_lb_) arises as a prolongation of the middle sternal
region in front of the mouth, while the bilobed metastoma is an
outgrowth of the sternal region behind it. {219}
The posterior cephalic and the thoracic appendages (_5–14_) elongate
and gradually approach the form which they possess in the adult. I
have not been able to discover, at any period of development, an
outer division or exopodite in any of the five posterior thoracic
limbs. And this is a very remarkable circumstance, inasmuch as such
an exopodite exists in the closely allied lobster in the larval
state; and, in many of the shrimp and prawn-like allies of the
crayfish, a complete or rudimentary exopodite is found in these
limbs, even in the adult condition.
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