The Crayfish: An Introduction to the Study of Zoology.Huxley, Thomas Henry
Science
The Crayfish: An Introduction to the Study of Zoology.
Huxley, Thomas Henry
Crayfish; Zoology
VII. 0 (ep.) 0 0 0 = 0
VIII. 1 1 0 0 = 2
IX. 1 1 1 0 = 3
X. 0 1 1 0 = 2
XI. 0 0 0 1 = 1
XII. 0 0 0 1 = 1
XIII. 0 0 0 0 = 0
XIV. 0 0 0 0 = 0
── ── ── ── ──
2 + ep. + 3 + 2 + 2 = 9 + ep.
It will be observed that the suppression of branchiæ has here taken
place in all the series, and at both the anterior and the posterior
ends of each. But the defect in total number is made up by the
increase of size, not of the pleurobranchiæ alone, as in the case
of the prawns, but of the arthrobranchiæ as well. At the same time
the whole apparatus has become more specialized and perfected as a
breathing organ. The close fitting of the edges of the carapace,
and the possibility of closing the inhalent and exhalent apertures,
render the crabs much more independent of actual immersion in water
than most of their congeners; and some of them habitually live on dry
land and breathe by means of the atmospheric air which they take into
and expel from their branchial cavities.
Notwithstanding all these wide departures from the structure and
habits of the crayfishes, however, attentive examination shows that
the plan of construction of the {278} crab is, in all fundamental
respects, the same as that of the crayfish. The body is made up of
the same number of somites. The appendages of the head and of the
thorax are identical in number, in function, and even in the general
pattern of their structure. But two pairs of abdominal appendages
in the female, and four pairs in the male, have disappeared. The
exopodites of the antennæ have vanished, and not even epipodites
remain to represent the podobranchiæ of the posterior five pairs
of thoracic limbs. The exceedingly elongated eye-stalks are turned
backwards and outwards, above the bases of the antennules and
the antennæ, and the bases of the latter have become united with
the edges of the carapace in front of them. In this manner the
extraordinary face, or _metope_ (fig. 72, B) of the crab results from
a simple modification of the arrangement of parts, every one of which
exists in the crayfish. The same common plan serves for both.
* * * * *
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